Origin of Species Variorum

↑ 1 blocks not present in 1859 1860 1861 1866 1869; present in 1872

Breeders of cattle wish the flesh and fat to be well marbled together: an animal thus characterised has been slaughtered, but the breeder has gone with confidence to the same stock and has succeeded.

→ a well-flavoured vegetable is cooked, and the individual is destroyed; 1859 1860 1861

breeders of cattle wish the flesh and fat to be well marbled together; the animal has been slaughtered, 1866 1869

→ horticulturist sows seeds of 1859 1860 1861

breeder has gone with confidence to 1866 1869

→ confidently expects to get nearly the same variety: breeders of cattle wish the flesh and fat to be well marbled together; the animal has been slaughtered, but the breeder goes with confidence to the same family. 1860

confidently expects to get nearly the same variety; breeders of cattle wish the flesh and fat to be well marbled together; the animal has been slaughtered, but the breeder goes with confidence to the same family. 1859

confidently expects to get nearly the same variety: breeders of cattle wish the flesh and fat to be well marbled together; the animal has been slaughtered, but the breeder goes with confidence to the same stock. 1861

has succeeded. 1866 1869

→ in 1859 1860 1861

may be placed in 1866 1869 1872

→ I do not doubt that 1859 1860 1861

probably 1866 1869

OMIT 1872

→ be slowly 1859 1860 1861 1866 1869

it is probable, be 1872

→ Thus I believe it has been with social insects: a 1859 1860 1861 1866 1869

Hence we may conclude that 1872

→ has been advantageous to the community: 1859 1860 1861 1866 1869

have proved advantageous: 1872

→ of the same community 1859 1860 1861 1866 1869

have 1872

→ And I believe that 1859 1860 1861 1866 1869

OMIT 1872

→ until 1859 1860 1861 1866 1869

many times, until 1872

desired end. ↑ Thus, →a well-flavoured vegetable is cooked, and the individual is destroyed; but the →horticulturist sows seeds of the same

family,

stock

stock,

and →confidently expects to get nearly the same variety: breeders of cattle wish the flesh and fat to be well marbled together; the animal has been slaughtered, but the breeder goes with confidence to the same family.

~~I have~~

I have

Such

such

faith →in the

power

powers

of selection, that →I do not doubt that a breed of cattle, always yielding oxen with extraordinarily long horns,

could,

could

→be slowly formed by carefully watching which individual bulls and cows, when matched, produced oxen with the longest horns; and yet no one ox

would

could

ever have propagated its kind. →Thus I believe it has been with social insects: a slight

modifications

modification

structure

structure,

of instinct,

instinct,

correlated with the sterile condition of certain members of the community, →has been advantageous to the community: consequently the fertile males and females →of the same community flourished, and transmitted to their fertile offspring a tendency to produce sterile members

with

having

the same

modifications.

modification.

→And I believe that

This

this

process

must have

has

been

repeated

repeated,

→until that prodigious amount of difference between the fertile and sterile females of the same species has been produced, which we see in

so many

many

social insects.

But we have not as yet touched on the climax of the difficulty; namely, the fact that the neuters of several ants differ, not only from the fertile females and males, but from each other, sometimes to an almost incredible degree, and are thus divided into two or even three castes. The castes, moreover, do not

commonly

generally

graduate into each other, but are perfectly well defined; being as distinct from each

other

other,

as are any two species of the same genus, or rather as any two genera of the same family. Thus in Eciton, there are working and soldier neuters, with jaws and instincts extraordinarily