Origin of Species Variorum

→ the colts have 1859 1860 1861 1866

this is 1869 1872

→ acquired their full amount of proportional difference. 1859 1860 1861 1866

the case. 1869 1872

→ the evidence appears to me conclusive, 1859 1860 1861 1866

we have conclusive evidence 1869 1872

→ Pigeon have 1859 1860 1861 1866

the Pigeon are 1869 1872

→ young pigeons of various breeds, 1859 1860 1861 1866

the young 1869 1872

→ the 1859 1860 1861 1866 1872

in the wild parent-species, in pouters, fantails, runts, barbs, dragons, carriers, and tumblers, the 1869

→ so extraordinarily in 1859 1860 1861 1866

in so extraordinary a degree in the 1869

in so extraordinary a manner in the 1872

→ that 1859 1860 1861 1866

and in other characters, that 1869 1872

→ I cannot doubt, be ranked in 1859 1860 1861

certainly have been ranked as 1866 1872

certainly be ranked as 1869

→ had they been natural productions. 1859 1860 1861 1866

if found in a state of nature. 1869 1872

→ be distinguished from each other, yet their 1859 1860 1861 1866

just be distinguished, yet the 1869

just be distinguished, the 1872

→ from 1859 1860 1861 1866 1872

in all its proportions from 1869

→ in all its proportions, 1859 1860 1861 1866

in 1872

OMIT 1869

→ as much 1859 1860 1861 1866 1869

the same proportions 1872

→ The 1859 1860 1861 1866 1869

These facts are explained by the above 1872

→ two principles above given seem to me to 1859 1860 1861

two principles above given, namely that variations do not generally supervene at a very early age, and that they are inherited at a corresponding age whatever that may have been, seem to me to 1866

above two principles 1869

two principles. 1872

→ facts in regard to the later embryonic stages of our domestic varieties. Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been acquired earlier or later in life, if the full-grown animal possesses them. 1859 1860 1861

OMIT 1872

several facts regarding the later developmental stages of our domestic varieties. 1866

facts. Fanciers select for breeding their dogs, horses, pigeons, &c., when nearly grown up: they are indifferent whether the desired qualities are acquired earlier or later in life, if the full-grown animal possesses them. 1869

but having had careful measurements made of the

dams

dam

and of

the

three-days-old

three-days

~~old~~

old

colts

colt

race-horses

race

a race

and heavy

cart-horses,

cart-horse,

I find that →the colts have by no means →acquired their full amount of proportional difference.

As →the evidence appears to me conclusive, that the

~~several domestic~~

several domestic

breeds of →Pigeon have descended from

a single

one

wild species, I compared →young pigeons of various breeds, within twelve hours after being hatched; I carefully measured →the proportions (but will not here give

the details)

details)

of the beak, width of mouth, length of nostril and of eyelid, size of feet and length of

leg.

leg,

~~the~~

the

~~wild~~

wild

parent-species,

~~stock,~~

~~pouters,~~

~~fantails,~~

~~runts,~~

~~barbs,~~

~~dragons,~~

~~carriers,~~

~~and~~

~~tumblers.~~

Now some of these birds, when mature, differ →so extraordinarily in length and form of beak, →that they

would

would,

→I cannot doubt, be ranked in distinct

genera

genera,

→had they been natural productions. But when the nestling birds of these several breeds were placed in a row, though most of them could →be distinguished from each other, yet their proportional differences in the above specified

~~several~~

several

points were incomparably less than in the full-grown birds. Some characteristic points of difference— for instance, that of the width of mouth— could hardly be detected in the young. But there was one remarkable exception to this rule, for the young of the short-faced tumbler differed →from the young of the wild rock-pigeon and of the other breeds, →in all its proportions, almost exactly →as much as in the adult state.

→The →two principles above given seem to me to

~~explain~~

explain

~~these~~

these

→facts in regard to the later embryonic stages of our domestic varieties. Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been acquired earlier or later in life, if the full-grown animal possesses them.