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1859
1861
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1869
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respects. 1860 1861 1866 1872
respects in the individuals of the same species. 1869

the degree 1860 1861 1866 1869
also, the extent 1872

in certain groups. 1859 1860 1861 1866 1869
belonging to the same family. 1872

we find in an 1859 1860 1861 1866
in certain 1869 1872

no trace of an organ, 1859 1860 1861 1866
parts are entirely absent 1869 1872

and which is 1859 1860 1861 1866
and which are 1869
in them, and which are 1872

the snapdragon (antirrhinum) we generally do not find a rudiment of a 1859 1860 1861
some Scrophulariaceæ we rarely find even a rudiment of a 1866
most of the Scrophulariaceæ the 1869 1872

but this may sometimes 1859 1860 1861 1866
is utterly aborted; yet we may conclude that a fifth stamen once existed, for a rudiment of it is found in many species of the family, and this rudiment occasionally becomes perfectly developed, as may 1869
is utterly aborted; yet we may conclude that a fifth stamen once existed, for a rudiment of it is found in many species of the family, and this rudiment occasionally becomes perfectly developed, as may sometimes 1872

seen. 1859 1860 1861
seen plainly or fully developed. 1866
seen in the common snap-dragon. 1869 1872

or more necessary, 1859 1860 1861 1866 1869
or, in order fully to understand the relations of the parts, more useful 1872

than the use and 1859 1860 1861 1866
in order fully to understand the relations of the parts, than the 1869
than the 1872

leg of the 1859 1860 1861 1866 1869
OMIT 1872

relatively to the adjoining parts in the embryo, 1859 1860 1861 1866 1869
in the embryo relatively to the adjoining parts, 1872

and in other respects.
In
Moreover, in
closely allied species, the degree to which the same organ has been
reduced
rendered rudimentary
occasionally differs much. This latter fact is well exemplified in the state of the wings
of
in
of the
female moths in certain groups. Rudimentary organs may be utterly aborted; and this implies, that we find in an
animals
animal
or
plants,
plant
no trace of an organ, which analogy would lead us to expect to
find
find,
and which is occasionally found in monstrous
individuals.
individuals
of
of
the
the
species.
species.
Thus in the snapdragon (antirrhinum) we generally do not find a rudiment of a fifth
stamen
stamen;
but this may sometimes be seen. In tracing the homologies of
any
the same
part in different
mem- bers
members
of
the same
a
class, nothing is more common, or more necessary, than the use and discovery of rudiments. This is well shown in the drawings given by Owen of the
leg-bones
bones
of the leg of the horse, ox, and rhinoceros.
It is an important fact that rudimentary organs, such as teeth in the upper jaws of whales and ruminants, can often be detected in the embryo, but afterwards wholly disappear. It is also, I believe, a universal rule, that a rudimentary part
or organ
or organ
is of greater size relatively to the adjoining parts in the embryo, than in the adult; so that the organ at this early age is less rudimentary, or even cannot be said to be in any degree rudimentary.
Hence
Hence,
also, a
also, a
rudimentary
organs
organ
in the
adult,
adult
are
is
often said to have retained
their
its
embryonic condition.
I have now given the leading facts with respect to rudimentary organs. In reflecting on them, every one must be struck with
astonishment;
astonishment:
for the same reasoning power which tells us
plainly
plainly
that most parts and organs are exquisitely adapted for certain purposes, tells us with equal plainness that these rudimentary or atrophied
organs
organs,
are imperfect and useless. In works