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1859
1860
1861
1866
1869
1872

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1859
1861
1866
1869
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reduced, during successive generations, 1859 1860 1866 1869
reduced, during successive genera- tions, 1861
formerly reduced 1872

having been better fitted by 1860
having been fitted by 1859
or lips, having become better fitted by 1861 1866 1869
or lips, having become excellently fitted through 1872

untouched by selection or disuse, 1859 1860 1861 1866 1869
unaffected, 1872

being and each separate organ 1859 1860 1861
being and each of its separate parts 1866
being with all its separate parts 1869
with all its separate parts 1872

parts, like 1859 1860 1861
organs, like 1866
organs bearing the plain stamp of inutility, such as 1869 1872

bear the plain stamp of inutility! 1860 1861 1866
bear the plain stamp of inutility! Nature may be said to have taken pains to reveal, by rudimentary organs and by homologous structures, her scheme of modification, which it seems that we wilfully will not understand. 1859
occur. 1869 1872

by rudimentary organs and by homologous structures, 1860 1861 1866
OMIT 1869 1872

which it seems that we wilfully will not understand. 1860 1861 1866
by means of rudimentary organs, embryological and homologous structures, but we wilfully will not understand the scheme. 1869
by means of rudimentary organs, of em- bryological and homologous structures, but we are too blind to understand her meaning. 1872

been modified, during a long course of descent, by the preservation or the natural selection of many successive slight favourable variations. 1860 1861 1866
changed, and are still slowly changing by the preservation and accumulation of successive slight favourable variations. 1859
been modified, during a long course of descent, chiefly through the natural selection of numerous successive, slight, favourable variations. 1869
been modified, during a long course of descent. 1872

8 blocks not present in 1859 1860 1861 1866 1869; present in 1872
This has been effected chiefly through the natural selection of numerous successive, slight, favourable variations; aided in an important manner by the inherited effects of the use and disuse of parts; and in an unimportant manner, that is in relation to adaptive structures, whether past or present, by the direct action of external conditions, and by variations which seem to us in our ignorance to arise spontaneously. It appears that I formerly underrated the frequency and value of these latter forms of variation, as leading to permanent modifications of structure independently of natural selection. But as my conclusions have lately been much misrepresented, and it has been stated that I attribute the modification of species exclusively to natural selection, I may be permitted to remark that in the first edition of this work, and subsequently, I placed in a most conspicuous position— namely, at the close of the Introduction— the following words: " I am convinced that natural selection has been the main but not the exclusive means of modification." This has been of no avail. Great is the power of steady misrepresentation; but the history of science shows that fortunately this power does not long endure. It can hardly be supposed that a false theory would explain, in so satisfactory a manner as does the theory of natural selection, the several large classes of facts above specified. It has recently been objected that this is an unsafe method of arguing; but it is a method used in judging of the common events of life, and has often been used by the greatest natural philosophers. The undulatory theory of light has thus been arrived at; and the belief in the revolution of the earth on its own axis was until lately supported by hardly any direct evidence.

of life; and we can
clearly
clearly
understand on this view the meaning of rudimentary organs. But disuse and selection will generally act on each creature, when it has come to maturity and has to play its full part in the struggle for existence, and will thus have little power
of acting
of acting
on an organ during early life; hence the organ will not be
much
much
reduced or rendered rudimentary at this early age. The calf, for instance, has inherited teeth, which never cut through the gums of the upper jaw, from an early progenitor having well-developed teeth; and we may believe, that the teeth in the mature animal were reduced, during successive generations, by
disuse,
disuse
owing to
or by
the tongue and
palate,
palate
having been better fitted by natural selection to browse without their aid; whereas in the calf, the teeth have been left untouched by selection or disuse, and on the principle of inheritance at corresponding ages have been inherited from a remote period to the present day. On the view of each
organism
organic
being and each separate organ having been specially created, how utterly
in-explicable
inexplicable
is it
it is
that parts, like the teeth in the embryonic calf or
like
like
the shrivelled wings under the soldered wing-covers of
many
some
beetles, should
thus
thus
so frequently bear the plain stamp of inutility! Nature may be said to have taken pains to
reveal
reveal,
by rudimentary organs and by homologous structures, her scheme of modification, which it seems that we wilfully will not understand.
I have now recapitulated the
chief
chief
facts and considerations which have thoroughly convinced me that species have been modified, during a long course of descent, by the preservation or the natural selection of many successive slight favourable variations. I cannot believe that a false theory would explain, as it seems to me that the theory of natural selection does explain,