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this process of the continued preservation or natural selection of more and more attractive flowers, 1859 1860 1861 1866
the above process long continued, 1869 1872

can most effectually 1859 1860 1861
can effectually 1866
OMIT 1869 1872

one— not as a very striking case, but 1859 1860 1861 1866
one, 1869 1872

pollen-grains, 1859 1860 1866
pollen grains, 1861
a few pollen-grains, 1869
a few pollen grains, 1872

accidentally dusted with pollen, having 1859 1860 1861 1866
which had 1869 1872

When our plant, by this process of the continued preservation or natural selection of more and more attractive flowers, had been rendered highly attractive to insects, they would, unintentionally on their part, regularly carry pollen from flower to flower; and that they can most effectually do
this
this,
effectually, I
I
could easily show by many striking
facts.
instances.
I will give only one— not as a very striking case, but as likewise illustrating one step in the separation of the sexes of
plants.
plants,
presently
presently
to
to
be
be
alluded
alluded
to.
to.
Some holly-trees bear only male flowers, which have four stamens producing
rather a
a rather
small quantity of pollen, and a rudimentary pistil; other
holly trees
holly-trees
bear only female flowers; these have a full-sized pistil, and four stamens with shrivelled anthers, in which not a grain of pollen can be detected. Having found a female tree exactly sixty yards from a male tree, I put the stigmas of twenty flowers, taken from different branches, under the microscope, and on all, without exception, there were pollen-grains, and on some a
profusion.
profusion
of
of
pollen.
pollen.
As the wind had set for several days from the female to the male tree, the pollen could not thus have been carried. The weather had been cold and boisterous, and therefore not favourable to bees, nevertheless every female flower which I examined had been effectually fertilised by the bees, accidentally dusted with pollen, having flown from tree to tree in search of nectar. But to return to our imaginary case: as soon as the plant had been rendered so highly attractive to insects that pollen was regularly carried from flower to flower, another process might commence. No naturalist doubts the advantage of what has been called the "physiological division of labour;" hence we may believe that it would be advantageous to a plant to produce stamens alone in one flower or on one whole plant, and pistils alone in