Origin of Species Variorum

→ (utterly ignorant though we be of the meaning of the law) 1859 1860 1861 1866

OMIT 1869 1872

→ intervals— 1859 1860 1861 1866

intervals of time— 1869 1872

↑ 1 blocks not present in 1859 1860 1861 1866; present in 1869 1872

notwithstanding that the plants own anthers and pistil stand so near each other as almost to ensure self-fertilisation, the fullest freedom for the entrance of pollen from another individual will explain the above state of exposure of the organs.

→ in several, perhaps in all, such flowers, there is a very 1859 1860

in most of these flowers there is a very 1861 1866

in most of these flowers there is a 1869

these almost invariably present beautiful and 1872

→ between the structure of the flower and the manner in which bees suck the nectar; for, in doing this, they either push the flower's own pollen on the stigma, or bring pollen from another flower. 1859 1860

between their structure and the manner in which bees suck the nectar; for, in doing this, they either push the flower's own pollen on the stigma, or bring pollen from another flower. 1861 1866

between their structure and the manner in which bees suck the nectar; for, in doing this, they either push the flowers own pollen on the stigma, or bring pollen from another flower. 1869

in relation to the visits of insects. 1872

→ I have found, by experiments published elsewhere, that 1859 1860 1861 1866 1869

OMIT 1872

→ that bees should 1859 1860 1861 1866 1869

for insects to 1872

→ as I believe, 1859 1860 1861 1866 1869

OMIT 1872

→ touch 1859 1860 1861 1866 1869

ensure fertilisation, just to touch with the same brush 1872

fertility to the offspring; and on the other hand, that close interbreeding diminishes vigour and fertility; that these facts alone incline me to believe that it is a general law of nature →(utterly ignorant though we be of the meaning of the law) that no organic being

fertilises

self-fertilises

itself for

a perpetuity

an eternity

of generations; but that a cross with another individual is occasionally— perhaps at

~~very~~

very

long →intervals— indispensable.

On the belief that this is a law of nature, we can, I think, understand several large classes of facts, such as the following, which on any other view are inexplicable. Every hybridizer knows how unfavourable exposure to wet is to the fertilisation of a flower, yet what a multitude of flowers have their anthers and stigmas fully exposed to the weather!

but if

an occasional cross be indispensable, the fullest freedom for the entrance of pollen from another individual will explain this state of exposure, more especially as the plant's own anthers and pistil generally stand so close together that self-fertilisation seems almost inevitable. ↑ Many flowers, on the other hand, have their organs of fructification closely enclosed, as in the great papilionaceous or pea-family; but →in several, perhaps in all, such flowers, there is a very curious

adaptations

adaptation

→between the structure of the flower and the manner in which bees suck the nectar; for, in doing this, they either push the flower's own pollen on the stigma, or bring pollen from another flower. So necessary are the visits of bees to

many papilionaceous

many papilinaceous

papilionaceous

flowers, that →I have found, by experiments published elsewhere, that their fertility is greatly diminished if these visits be prevented. Now, it is scarcely possible →that bees should fly from flower to flower, and not

to carry

carry

pollen from one to the other, to the great

good

good,

→as I believe, of the plant.

Insects

Bees will

act like a camel-hair pencil, and it is

~~quite~~

quite

sufficient,

sufficient

~~just~~

just

to →touch the anthers of