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1859
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1861
1866
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1866
1869
1872

and in nearly every class, 1861 1866
OMIT 1869 1872

favourable variations 1861 1866
variations or individual differences of a favourable nature 1869
variations or individual differences of a favorable nature 1872

A difficulty, diametrically opposite to this which we have just been considering, has been advanced, namely, 1861 1866
OMIT 1869 1872

could 1861 1866
it has been asked, could 1869 1872

advancement or in the differentiation and specialisation 1861 1866
the advancement or differentiation 1869 1872

2 blocks not present in 1859 1860 1861; present in 1866 1869 1872
Mr. Herbert Spencer would probably answer that as soon as the most simple unicellular organism came by growth or division to be compounded of several cells, or became attached to any supporting surface, his law would come into action, namely, "that homologous units of any order become differentiated in proportion as their relations to incident forces become different." But as we have no facts to guide us, all speculation on the subject is useless.

through their preservation either 1861
through their preservation, either 1866
thus 1869 1872

I must recur to what was stated 1861
as I remarked 1866 1869 1872

where I say that 1861
OMIT 1866 1869 1872

due allowance be made 1861 1866
we make due allowance 1869 1872

many past epochs in its history. 1861 1866
past epochs in its history. 1869
present time, and still more so during past ages. 1872

exist
in numbers
in numbers
throughout the world, and in nearly every class, from various causes. In some cases favourable variations may never have arisen for natural
selec- tion
selection
to act on and accumulate. In no case, probably, has time sufficed for the utmost possible amount of development. In some few cases there
has
may have
been what we must call retrogression of organisation. But the main cause lies in the
fact
circumstance
that under very simple conditions of life a high organisation would be of no service,— possibly would be of actual disservice, as being of a more delicate nature, and more liable to be put out of order and
thus
thus
injured.
A difficulty, diametrically opposite to this which we have just been considering, has been advanced, namely,
that, looking
Looking
looking
to the
first dawn
dawn
of life, when all organic beings, as we may
believe,
imagine,
presented the simplest structure,
how,
how
could the first steps in advancement or in the differentiation and specialisation of parts have arisen? I can make no sufficient answer; and can only say that as we have no facts to guide us, all speculation on the subject would be baseless and useless. It is, however, an error to suppose that there would be no struggle for existence, and, consequently, no natural selection, until many forms had been produced: variations in a single species inhabiting an isolated station might be beneficial,
and,
and
through their preservation either the whole mass of individuals might
be
become
modified, or two distinct forms might arise.
But,
But
I must recur to what was stated towards the close of the Introduction, where I say that no one ought to feel surprise at much remaining as yet unexplained on the origin of species, if due allowance be made for our profound ignorance on the mutual relations of the inhabitants of the world
at
during
the many past epochs in its history.