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modification. 1859 1860 1861 1866
modification may always be expected. 1869 1872

these variable characters, produced by man's selection, sometimes become attached, 1859 1860 1861 1866
characters, modified through selection by man, are sometimes transmitted, 1869

much variability in the
parts
structure
undergoing modification. It further deserves notice that these variable characters, produced by man's selection, sometimes become attached, from causes quite unknown to us, more to one sex than to the other, generally to the male sex, as with the wattle of carriers and the enlarged crop of pouters.
Now let us turn to nature. When a part has been developed in an extraordinary manner in any one species, compared with the other species of the same genus, we may conclude that this part has undergone an extraordinary amount of
modification,
modification
since the period when the
several species
species
branched off from the common progenitor of the genus. This period will seldom be remote in any extreme degree, as species
very
very
rarely endure for more than one geological period. An extraordinary amount of modification implies an unusually large and long-continued amount of variability, which has continually been accumulated by natural selection for the benefit of the species. But as the variability of the
extraordinarily-developed
extraordinarily developed
part or organ has been so great and long-continued within a period not
exces- sively
excessively
remote, we might, as a general rule,
still expect
expect still
to find more variability in such parts than in other parts of the
organisation,
organisation
which have remained for a much longer period nearly constant. And this, I am convinced, is the case. That the struggle between natural selection on the one hand, and the tendency to reversion and variability on the other hand, will in the course of time cease; and that the most abnormally developed organs may be made constant, I
can
can
see no reason to doubt.
Hence,
Hence
when an organ, however abnormal it may be, has been transmitted in approximately the same condition to many modified descendants, as in the case of the wing of the bat, it must