| → we cannot doubt that 1861 1866 | 
| I have no doubt that 1859 1860 | 
| OMIT 1869 1872 | 
  | 
| → due to some quite distinct cause, probably 1859 1860 1861 1866 | 
| probably in chief part due 1869 1872 | 
  | 
| → the hooks 1859 1860 1861 1866 | 
| and which there is reason to believe from the distribution of the thorn-bearing species in Africa and South America, serves as a defence against browsing quad- rupeds, so the hooks 1869 | 
| and which, as there is reason to believe from the distribution of the thorn-bearing species in Africa and South America, serve as a defence against browsing quadrupeds, so the spikes 1872 | 
  | 
| → have arisen from unknown laws of growth, and 1859 1860 1861 1866 | 
| first 1869 | 
| at first 1872 | 
  | 
| → subsequently 1859 1860 1861 1866 | 
| developed for this object, and subsequently been 1869 | 
| developed for this object, and subsequently have been improved and 1872 | 
  | 
| → undergoing 1859 1860 1861 1866 | 
| as it underwent 1869 1872 | 
  | 
| → producing slight and unimportant variations; 1859 1860 1861 1866 | 
| of each slight variation or individual difference; 1869 1872 | 
  | 
| ↑ 1 blocks not present in  1859 1860 1861 1866 1869; present in  1872 | 
|  With cattle susceptibility to the attacks of flies is correlated with colour, as is the liability to be poisoned by certain plants; so that even colour would be thus subjected to the action of natural selection. 
 | 
  | 
| → on the whole organisation 1861 1866 1869 | 
| OMIT 1872 | 
  | 
| → is 1861 1866 1869 | 
| on the whole organisation is 1872 | 
  |