Origin of Species Variorum

→ in accordance with their diverse 1859 1860 1861 1866

for different 1869 1872

→ at a rather late 1859 1860 1861 1866

in the several species at a not early 1869 1872

→ of the new species of our supposed genus will manifestly tend to 1859 1860 1861 1866

will be left but little modified and will 1869

will have been but little modified, and they will still 1872

→ whole families or even 1859 1860 1861 1866

widely distinct structures and to whole 1869 1872

→ in the parent-species, may have become, by 1860 1861 1866

in the parent-species, may become, by 1859

to a remote progenitor, may have become, through 1869 1872

→ principles— namely of each successive modification supervening at a rather late age, and being inherited at a corresponding late age— 1859 1861 1866

principles— namely of each successive modification supervening at a rather late age, and being inherited at a corre- sponding late age— 1860

principles 1869 1872

→ in 1859 1860 1861 1866

will not have been much modified in 1869 1872

→ descendants of the parent species will still resemble each other closely, for they will not have been modified. 1861

descendants of the parent-species will still resemble each other closely, for they will not have been modified. 1859 1860 1866

forms; although in each the embryonic fore-limb will differ greatly from that in the adult. 1869

forms; although in each form the fore-limb will differ greatly in the adult state. 1872

→ long-continued exercise or use on the one hand, and disuse on the other, 1859 1860 1861

long-continued exercise or disuse 1866

moreover, long-continued use or disuse 1869

long-continued use or disuse 1872

→ in modifying an organ, such influence will mainly affect the mature animal, which has come to its 1859 1860 1861

had in modifying an organ, such influence will mainly affect the mature animal, which has come to its 1866

had in modifying the limbs or other parts of animals, this will chiefly or solely have affected them when mature and when they had to use their 1869

had in modifying the limbs or other parts of any species, this will chiefly or solely have affected it when nearly mature, when it was compelled to use its 1872

→ of activity and has 1859 1860 1861 1866

OMIT 1869 1872

→ be inherited 1859 1860 1861 1866

be transmitted to the offspring 1869

have been transmitted to the offspring 1872

→ remain unmodified, or 1859 1860 1861 1866

not 1869

not be modified, or 1872

→ be modified 1859 1860 1861 1866 1869

will be modified only 1872

→ in a lesser degree, by the effects of use and disuse. 1859 1860 1861 1866

or will be modified in a less degree. 1869

in a slight degree, through the effects of the increased use or disuse of parts. 1872

→ In certain cases 1859 1860 1861 1866

In other cases 1869

With some animals 1872

→ steps of variation might supervene, from causes of which we are wholly ignorant, 1859 1860 1861 1866

variations may have supervened 1869 1872

modified through natural selection →in accordance with their diverse habits. Then, from the many slight successive

~~steps of~~

steps of

variations

variation

having supervened →at a rather late age, and having been inherited at a corresponding age, the young →of the new species of our supposed genus will manifestly tend to resemble each other much more closely than do the

adults,—

adults,

just as we have seen

with

the

breeds

case

the pigeon.

pigeons.

We may extend this view to →whole families or even classes. The fore-limbs, for instance, which

once served

served

as legs →in the parent-species, may have become, by a long course of modification, adapted in one descendant to act as hands, in another as paddles, in another as wings;

but

and

on the above two →principles— namely of each successive modification supervening at a rather late age, and being inherited at a corresponding late age— the fore-limbs →in the embryos of

these

the

several →descendants of the parent species will still resemble each other closely, for they will not have been modified. But in each

individual

of our

new species, the embryonic fore-limbs will differ greatly from the fore-limbs in the mature animal; the limbs in the

later

latter

having undergone much modification at a rather late period of life, and having thus been converted into hands, or paddles, or wings. Whatever

influence,

influence

→long-continued exercise or use on the one hand, and disuse on the other, may have →in modifying an organ, such influence will mainly affect the mature animal, which has come to its full powers →of activity and has

had to

gain

their

its

own living; and the

effect

effects

thus produced will →be inherited at

the

corresponding

nearly mature

mature