Origin of Species Variorum

→ all species have 1859 1860 1861 1866

species have all 1869 1872

→ which my theory requires, 1859 1860 1861 1866

required, 1869

required by the theory, 1872

→ formations distant from each other in time. 1859 1860 1861 1866

widely separated formations. 1869 1872

→ can 1859 1860 1861 1866 1869

as far as I can see, may 1872

→ do not 1859 1860 1861 1866

are not 1869

are by no means sufficient to 1872

→ overthrow 1859 1860 1861 1866 1872

sufficient to overthrow 1869

→ from a few primordial forms with subsequent 1861 1866

with 1859 1869 1872

from a few created forms with subsequent 1860

→ variability. 1859 1860 1861 1866

variability, caused, or at least excited, by changed conditions of life. 1869

variability, caused, or at least excited, by changed conditions of life; but often in so obscure a manner, that we are tempted to consider the variations as spontaneous. 1872

→ by 1859 1860 1861 1866 1869

compensation, the increased 1872

→ and by the direct 1859 1860 1861 1866

and by the definite 1869

of parts, and the definite 1872

of time, geology plainly declares that →all species have changed; and they have changed in the manner →which my theory requires, for they have changed slowly and in a graduated manner. We clearly see this in the fossil remains from consecutive formations invariably being much more closely related to each other, than are the fossils from →formations distant from each other in time.

Such is the sum of the several chief objections and difficulties which may

be justly

justly be

urged against

the

theory; and I have now briefly recapitulated the answers and explanations

which,

which

→can be

given.

given

~~them.~~

them.

I have felt these difficulties far too heavily during many years to doubt their weight. But it deserves especial notice that the more important objections relate to questions on which we are confessedly ignorant; nor do we know how ignorant we are. We do not know all the possible transitional gradations between the simplest and the most perfect organs; it cannot be pretended that we know all the varied means of Distribution during the long lapse of years, or that we know how imperfect

is the

the

Geological

Record.

Record

~~is.~~

is.

Serious

Grave

as these several

objections

difficulties

are, in my judgment they →do not →overthrow the theory of descent →from a few primordial forms with subsequent

subsequent modification.

modification.

Now let us turn to the other side of the argument. Under domestication we see much →variability. This seems to be

in part

mainly

due to the reproductive system being eminently susceptible to changes in the conditions of life; so that this system, when not rendered impotent, fails to reproduce offspring exactly like the parent-form.

This variability

Variability

is governed by many complex laws,— by

correlation,

correlated

correlation

growth,

~~of growth,~~

of growth,

→by use and

disuse

disuse,