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can most effectually 1859 1860 1861
can effectually 1866
OMIT 1869 1872

one— not as a very striking case, but 1859 1860 1861 1866
one, 1869 1872

pollen grains, 1861
pollen-grains, 1859 1860 1866
a few pollen-grains, 1869
a few pollen grains, 1872

accidentally dusted with pollen, having 1859 1860 1861 1866
which had 1869 1872

regularly carry pollen from flower to flower; and that they can most effectually do
this
this,
effectually, I
I
could easily show by many striking
facts.
instances.
I will give only one— not as a very striking case, but as likewise illustrating one step in the separation of the sexes of
plants.
plants,
presently
presently
to
to
be
be
alluded
alluded
to.
to.
Some holly-trees bear only male flowers, which have four stamens producing
rather a
a rather
small quantity of pollen, and a rudimentary pistil; other
holly trees
holly-trees
bear only female flowers; these have a full-sized pistil, and four stamens with shrivelled anthers, in which not a grain of pollen can be detected. Having found a female tree exactly sixty yards from a male tree, I put the stigmas of twenty flowers, taken from different branches, under the microscope, and on all, without exception, there were pollen grains, and on some a
profusion.
profusion
of
of
pollen.
pollen.
As the wind had set for several days from the female to the male tree, the pollen could not thus have been carried. The weather had been cold and boisterous, and therefore not favourable to bees, nevertheless every female flower which I examined had been effectually fertilised by the bees, accidentally dusted with pollen, having flown from tree to tree in search of nectar. But to return to our imaginary case: as soon as the plant had been rendered so highly attractive to insects that pollen was regularly carried from flower to flower, another process might commence. No naturalist doubts the advantage of what has been called the "physiological division of labour;" hence we may believe that it would be advantageous to a plant to produce stamens alone in one flower or on one whole plant, and pistils alone in another flower or on another plant. In plants under culture and placed under new conditions of life, sometimes the male organs and sometimes the female organs become more or less impotent; now if we suppose this to