Origin of Species Variorum

→ OMIT 1866

to the extinct and unknown 1859 1860 1861

is to the extinct and unknown, 1869 1872

→ is very small 1866

is very small, 1859 1860 1861

OMIT 1869 1872

→ compared with the extinct and unknown forms, I 1866

I 1859 1860 1861 1869 1872

→ It certainly is true, that new organs very rarely or never suddenly appear in any class, as if created for some special purpose; as indeed is 1866

The truth of this remark is indeed 1859 1860

It certainly is not true, that new organs often appear suddenly in any class, as if created for some special purpose; as indeed is 1861

It certainly is true, that new organs appearing as if specially created for some purpose, rarely or never appear suddenly in any class; as indeed is 1869

It certainly is true, that new organs appearing as if created for some special purpose, rarely or never appear in any being;— as indeed is 1872

→ but somewhat exaggerated, canon 1861 1866 1869 1872

canon 1859

but somewhat exaggerated canon 1860

→ so much variety and so little novelty? 1866 1869

so? 1859 1860 1861

so much variety and so little real novelty? 1872

→ sudden leap, 1866 1869

leap, 1859 1860 1861

great and sudden leap, 1872

→ short and sure though slow 1866 1869

the shortest and slowest 1859 1860

short and slow 1861

short and sure, though slow 1872

→ Importance, as affected by Natural Selection. 1866

importance .— 1859 1860 1861

Importance, as affected by Natural Selection. 1869

Importance, as affected by Natural Selection . 1872

→ individuals with any favourable variation, 1859 1860 1861 1866

the fittest, 1869 1872

→ those with any unfavourable deviation of structure,—I have sometimes felt much difficulty in understanding the origin of simple parts, of which the importance does not seem sufficient to cause the preservation of successively varying individuals. 1866

those with any unfavourable deviation of structure,— I have sometimes felt much difficulty in understanding the origin of simple parts, of which the importance does not seem sufficient to cause the preservation of successively varying individuals. 1859 1860 1861

the less well fitted individuals,—I have sometimes felt great difficulty in understanding the origin or formation of parts of little importance; almost as great, though of a very different kind, as in the case of the most perfect and complex organs. 1869

the less well-fitted individuals, 1872

conjecture by what transitions

many

....

organ

organs

could

....

have arrived at

its

their

present state; yet, considering

how small

that

the proportion of living and known forms →OMIT →is very small →compared with the extinct and unknown forms, I have been astonished how rarely an organ can be named, towards which no transitional grade is known to lead. →It certainly is true, that new organs very rarely or never suddenly appear in any class, as if created for some special purpose; as indeed is shown by that

old

old,

→but somewhat exaggerated, canon in natural history of "Natura non facit saltum." We meet with this admission in the writings of almost every experienced naturalist;

or,

....

as Milne Edwards has well expressed it,

nature

Nature

is prodigal in variety, but niggard in innovation. Why, on the theory of Creation, should

this

there

be →so much variety and so little novelty? Why should all the parts and organs of many independent beings, each supposed to have been separately created for its proper place in nature, be so

invariably

commonly

linked together by graduated steps? Why should not Nature

have

....

taken

take

leap

sudden leap

from structure to structure? On the theory of natural selection, we can clearly understand why she should not; for natural selection

can

....

act

acts

only by taking advantage of slight successive variations; she can never take a →sudden leap, but must advance by →short and sure though slow steps.

Organs

little

apparent

→ Importance, as affected by Natural Selection.

As natural selection acts by life and

death,—

death,—by

....

the

survival

preservation

of →individuals with any favourable variation, and by the destruction of →those with any unfavourable deviation of structure,—I have sometimes felt much difficulty in understanding the origin of simple parts, of which the importance does not seem sufficient to cause the preservation of successively varying individuals.