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something analogous 1866
the same thing 1869 1872

a short-styled cowslip yields more seed when fertilised by the long-styled 1866
instance, the mid-styled form of Lythrum salicaria was illegitimately fertilised with the greatest ease by pollen from the longer stamens of the short-styled 1869 1872

less 1866
yielded many seeds; but the latter form did not yield a single 1869 1872

its own form, than does a long-styled cowslip when fertilised in the two corresponding methods. 1866
the longer stamens of the mid-styled form. 1869 1872

the 1866
and in others which might have been adduced, the 1869
and in others which might be added, the 1872

not corresponding in height with the pistil, differs much in degree, up to absolute and utter sterility; just in the same manner as occurs in crossing distinct species. As the degree of sterility in the latter case depends in an eminent degree on the conditions of life being more or less favourable, so I have found it with illegitimate unions. It is well known that if pollen of a distinct species be placed on the stigma of a flower, and its own pollen be afterwards, even after a considerable interval of time, placed on the same stigma, its action is so strongly prepotent that it generally annihilates the effect of the foreign pollen; so it is with the pollen of the several forms of the same species, for legitimate pollen is strongly prepotent over illegitimate pollen, when both are placed on the same stigma. I ascertained this by fertilising several flowers, first illegitimately, and twenty-four hours afterwards legitimately, with pollen taken from a peculiarly coloured variety, and all the seedlings were similarly coloured; this shows that the legitimate pollen, though applied twenty-four hours subsequently, had wholly destroyed or prevented the action of the previously applied illegitimate pollen. Again, as in making reciprocal crosses between the same two species, there is occasionally a great difference in the result, so something analogous occurs with
trimorphic
dimorphic
plants; for a short-styled cowslip yields more seed when fertilised by the long-styled form, and less seed when fertilised by its own form, than does a long-styled cowslip when fertilised in the two corresponding methods.
In all these
respects,
respects
the forms of the same undoubted species when illegitimately united behave in exactly the same manner as do two distinct species when crossed. This led me carefully to observe during four years many seedlings, raised from several illegitimate