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the formation of 1859 1860 1861 1866
OMIT 1869 1872

can be of any service 1859 1860 1861 1866
are beneficial 1869 1872

the excretion of precious 1859 1860 1861 1866
removing matter so precious as 1869 1872

sometimes 1859 1860 1861 1866
have been known to 1869 1872

have appeared, not from unknown laws of growth, but 1859 1860 1861 1866
have been developed 1869
are developed 1872

were formed 1859 1860 1861 1866
have been developed 1869 1872

simple. 1859 1860 1861 1866 1869
comparatively simple; and we can understand to a large extent the laws governing their imperfect development. 1872

breeds,— 1859 1860 1861 1866
breeds of sheep,— 1869
breeds of sheep, 1872

monsters. 1859 1860 1861 1866 1869
monsters; but I doubt whether any of these cases throw light on the origin of rudimentary organs in a state of nature, further than by showing that rudiments can be produced; for the balance of evidence clearly indicates that species under nature do not undergo great and abrupt changes. 1872

I doubt whether any of these cases throw light on the origin of rudimentary organs in a state of nature, further than by showing that rudiments can be produced; for I doubt whether species under nature ever undergo abrupt changes. 1859 1860 1861 1866 1869
we learn from the study of our domestic productions that the disuse of parts leads to their reduced size; and that the result is inherited. 1872

to 1859 1860 1861 1866
by beasts of prey to 1869

minute
papilla;
papilla,
which often
represent
represents
the pistil in male flowers, and which is formed
merely
....
of
cellular
mere cellular
tissue, can thus act? Can we suppose that the formation of rudimentary
teeth
teeth,
which are subsequently absorbed, can be of any service to the rapidly growing embryonic calf by the excretion of precious phosphate of lime? When a
mans
man's
fingers have been amputated, imperfect nails sometimes appear on the
stumps,
stumps:
and I
I
could as soon believe that these vestiges of nails have appeared, not from unknown laws of growth, but in order to excrete horny matter, as that the rudimentary nails on the fin of the manatee were formed for this
same purpose.
purpose.
On
the
my
view of descent with modification, the origin of rudimentary organs is simple. We have plenty of cases of rudimentary organs in our domestic productions,— as the stump of a tail in tailless breeds,— the vestige of an ear in earless breeds,— the reappearance of minute dangling horns in hornless breeds of cattle, more especially, according to Youatt, in young animals,— and the state of the whole flower in the cauliflower. We often see rudiments of various parts in monsters. But I doubt whether any of these cases throw light on the origin of rudimentary organs in a state of nature, further than by showing that rudiments can be produced; for I doubt whether species under nature ever undergo abrupt changes. I believe that disuse has been the main agency; that it has led in successive generations to the gradual reduction of various organs, until they have become rudimentary,— as in the case of the eyes of animals inhabiting dark caverns, and of the wings of birds inhabiting oceanic islands, which have seldom been forced to take flight, and have ultimately lost the power of flying. Again, an organ useful under certain conditions, might become injurious under others, as with the wings of beetles living on small and exposed