Origin of Species Variorum

→ between his doubtful forms, 1859 1860 1861 1866 1869

OMIT 1872

→ and to 1859 1860 1861 1866

or 1869

and then to 1872

→ and higher stage may be, in some cases, due merely to 1859 1860 1866

and higher stage may be; in some cases, due merely to 1861

may, in some cases, be the simple result of 1869

may, in many cases, be the simple result of 1872

→ in two different regions; but I have not much faith in this view; and I attribute the passage of a variety, from a state in which it differs very slightly from its parent to one in which it differs more, 1859 1860 1861 1866

but in most cases they may be attributed 1869

→ action 1859 1860 1861 1866

gradual accumulative action 1869

→ in accumulating (as will hereafter 1859 1860 1861 1866

as hereafter to 1869

→ differences of structure in certain definite directions. 1859 1860 1861 1866

on fluctuating variability. 1869

↑ 1 blocks not present in 1859 1860 1861 1866 1869; present in 1872

the nature of the organism and of the different physical conditions to which it has long been exposed; but with respect to the more important and adaptive characters, the passage from one stage of difference to another, may be safely attributed to the cumulative action of natural selection, hereafter to be explained, and to the effects of the increased use or disuse of parts.

→ Hence I believe a 1859 1860 1861 1866

Hence a 1869

A 1872

and the truth of this admission will often be disputed by other naturalists.

When

When,

~~moreover,~~

moreover,

he comes to study allied forms brought from countries not now continuous, in which case he

can hardly

cannot

hope to find

~~the~~

the

intermediate

links,

links

→between his doubtful forms, he will

be compelled

have

to trust almost entirely to analogy, and his difficulties

will

....

rise to a climax.

Certainly no clear line of demarcation has as yet been drawn between species and

sub-species—

sub-species—that

that

....

is, the forms which in the opinion of some naturalists come very near to, but do not quite arrive

at,

the rank of

species:

species;

or, again, between sub-species and well-marked varieties, or between lesser varieties and individual differences. These differences blend into each other

an insensible series; and a series impresses the mind with the idea of an actual passage.

Hence I look at individual differences, though of small interest to the systematist, as of

the highest

high

importance for us, as being the first

steps

step

towards such slight varieties as are barely

thought

though

worth recording in works on natural history. And I look at varieties which are in any degree more distinct and permanent, as steps

towards

leading to

strongly-marked

strongly marked

and

~~more~~

permanent varieties; and at

the

these

latter, as leading to

sub-species

sub-species,

→and to species. The

passages

passage

from one stage of difference to another →and higher stage may be, in some cases, due merely to the long-continued action of different physical

conditions;

conditions

→in two different regions; but I have not much faith in this view; and I attribute the passage of a variety, from a state in which it differs very slightly from its parent to one in which it differs more, to the →action of natural

selection,

selection

→in accumulating (as will hereafter be more fully

explained,

explained)

→differences of structure in certain definite directions. ↑ →Hence I believe a well-marked variety may

be justly

therefore be

called an incipient