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1859
1860
1861
1866
1869
1872

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1859
1860
1861
1866
1872

The result depends on there being 1869
Its action depends on there being 1859 1860 1861 1866
It can act only when there are 1872

polity of nature, 1859 1860 1861 1866 1869
natural polity of a district 1872

some of the inhabitants of the country under-going 1869
some of the inhabitants of the country undergoing 1859 1860 1861
some of the inhabitants of the country under- going 1866
the 1872

kind. 1859 1860 1861 1866 1869
of its existing inhabitants. 1872

2 blocks not present in 1859 1860 1861 1866 1869; present in 1872
The occurrence of such places will often depend on physical changes, which generally take place very slowly, and on the immigration of better adapted forms being prevented. As some few of the old inhabitants become modified, the mutual relations of others will often be disturbed; and this will create new places, ready to be filled up by better adapted forms; but all this will take place very slowly.

more or less 1869
in some slight degree 1872

OMIT 1869
it would often be long before 1872

better adapted to the then existing conditions, may not soon 1869
in various parts of the organisation might 1872

3 blocks not present in 1869 1872; present in 1859 1860 1861 1866
But the action of natural selection will probably still oftener depend on some of the inhabitants becoming slowly modified; the mutual relations of many of the other inhabitants being thus disturbed. Nothing can be effected, unless favourable variations occur, and variation itself is apparently always a very slow process. The process will often be greatly retarded by free intercrossing.

But 1869 1872
On the other hand, 1859 1860 1861 1866

in effecting changes, at 1869
often only at 1859 1860 1861
generally at only 1866
only at 1872

results of natural selection accord perfectly 1869
action of natural selection accords perfectly 1859 1860 1861 1866
results accord 1872

the most favoured or improved varieties will
be
have been
enabled to spread: there will
be
have been
much extinction of the less improved forms, and the relative proportional numbers of the various inhabitants of the
renewed
reunited
continent will again
be
have been
changed; and again there will
be
have been
a fair field for natural selection to improve still further the inhabitants, and thus
produce
to produce
new species.
That natural selection
will always
always
generally
....
act
acts
with extreme
slowness,
slowness
I fully admit. The result depends on there being places in the polity of nature, which can be better
occupied by
filled through
some of the inhabitants of the country under-going
modification
modifications
of some kind. The existence of such places will often depend on physical changes, which are generally very slow, and on the immigration of better adapted forms
having been
being
checked. But the effects of natural selection will probably still oftener depend on some few of the inhabitants becoming slowly modified; the mutual relations of the other inhabitants being thus disturbed. Although all the individuals of the same species differ more or less from each other, OMIT differences of the right
nature
nature,
better adapted to the then existing conditions, may not soon occur. The
result
results
would
will
often be greatly retarded by free
intercrossing.
inter-crossing.
Many will exclaim that these several causes are amply sufficient
wholly
....
to
stop
neutralise
the
action
power
of natural selection. I do not believe so. But I do believe that natural selection
will always
always
will generally
generally
act
acts
very
slowly,
slowly
in effecting changes, at long intervals of time, and
generally
only
on
only
....
a
very
....
few of the inhabitants of the same
region
region.
at
....
the
....
same
....
time.
....
I further
believe,
believe
that
this very
this
these
slow,
intermit- tent
inter- mittent
intermittent
results of natural selection accord perfectly
well
....
with what geology tells us of the rate and manner at which the inhabitants of
this
the
world have changed.
Slow though the process of selection may be, if feeble