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at the close of 1869
in 1861 1866

asks, how, on the principle of natural selection, can a variety live side by side with the parent-species? 1869
has advanced various good objections to my views, and other remarks in its favour. 1861 1866

17 blocks not present in 1859 1860 1869 1872; present in 1861 1866
Of the objections, some seem to me unimportant, some few are owing to misapprehension, and some are incidentally noticed in various parts of this volume. On the erroneous supposition that all the species of a region are believed by me to be changing at the same time, he justly asks how it is that all the forms of life do not present a fluctuating and inextricably confused body? but it is sufficient for us if some few forms at any one time are variable, and few will dispute that this is the case. He asks, how can it be on the principle of natural selection that a variety should live in abundance side by side with the parent species; for the variety during its formation is supposed to have supplanted the intermediate forms between itself and the parent species, and yet it has not supplanted the parent species itself, for both are supposed now to live side by side? If the variety and parent species have become fitted to slightly different habits of life, they might live together; though in the case of animals which freely cross and move about, varieties seem to be almost always confined to distinct localities. But is it the case that varieties of plants and of the lower animals are often found in abundance side by side with the parent forms? Laying aside the polymorphic species in which the innumerable variations that occur seem neither advantageous nor disadvantageous to the species, and have not been fixed; laying aside also temporary variations, such as albinism, &c., my impression is that varieties and the supposed parent species are generally found, inhabiting either distinct stations, high land or low land, dry or moist districts, or distinct regions. Again, Professor Bronn truly remarks, that distinct species do not differ from each other in single characters alone, but in many; and he asks, how it comes that natural selection should always have simultaneously affected many parts of the organisation? Probably the whole amount of difference has not been simultaneously effected; and the unknown laws of correlation will certainly account for, but not strictly explain, much simultaneous modification. Anyhow, we see in our domestic varieties the very same fact: though our domestic races may differ much in some one organ from the other races of the same species, yet the other parts of the organisation will always be found in some degree different. Professor Bronn likewise asks with striking effect how, for instance in the mouse or hare genus, natural selection will account for the several species (descended, I may remark, from a parent of unknown character) having longer or shorter tails, longer or shorter ears, and fur of different colours; how will it account for one species of plant having pointed and another species obtuse leaves? I can give no definite answer to such questions; but I might ask in return, were these differences, on the doctrine of independent creation, formed for no purpose? If of use, or if due to correlation of growth, they could assuredly be formed through the natural preservation of such useful or correlated variations. I believe in the doctrine of descent with modification, notwithstanding that this or that particular change of structure cannot be accounted for, because this doctrine groups together and explains, as we shall see in the latter chapters, many general phenomena of nature. A distinguished botanist, Mr. H. C. Watson, believes that I have overrated the importance of the principle of divergence of character (in which, however, he apparently believes), and that convergence of character, as it may be called, has likewise played a part. This is an intricate subject which need not be here discussed. I will only say that if two species of two closely allied genera produced a number of new and divergent species, I can believe that these new forms might sometimes approach each other so closely that they would for convenience sake be classed in the same new genus, and thus two genera would converge into one; but from the strength of the principle of inheritance, it seems hardly credible that the two groups of new species would not at least form two sections of the supposed new single genus.

But such ill-considered objections are hardly worth notice.
The celebrated palæontologist,
Professor
....
Bronn, at the close of his German translation of this work, asks, how, on the principle of natural selection, can a variety live side by side with the parent-species? If both have become fitted for slightly different habits of life or conditions, they might live together; though, in the case of animals which freely cross and wander much about, varieties seem to be almost always confined to distinct localities. But if we put on one side polymorphic species, in which the variability seems to be of a peculiar nature, and all mere temporary variations, such as size, albinism, &c., the more permanent varieties are generally found, as far as I can judge, inhabiting distinct stations, high land or low land, dry or moist districts, or distinct regions. Bronn also insists that distinct species never differ from each other only in single characters, but in many parts; and he asks, how it comes that natural selection should invariably have affected simultaneously many parts of the organisation? But there is not the least necessity for believing that all the parts have been simultaneously modified; they may have been gained one after the other, and from being transmitted together, they appear to us as if simultaneously formed. Correlation, however, will account for various parts changing, when any one part changes. We have evidence of this in our domestic races, which though they may differ greatly in some one selected character, always differ to a certain extent in other characters.
Bronn, again, asks how natural selection can account for differences between species, which appear to be of no service to these species, such as the length of the ears or tail, or the folds of the enamel in the teeth, of