Origin of Species Variorum

→ from 1859 1860 1861 1866 1869

by kicking its 1872

→ of the smaller 1859 1860 1861 1866 1869

OMIT 1872

→ natural selection increased in successive generations 1859 1860 1861 1866 1869

OMIT 1872

→ its 1859 1860 1861 1866 1869

were increased during successive generations, its 1872

→ not quite decisive; 1866 1869

very scanty; 1861

not decisive; 1872

→ of inherited epilepsy in guinea-pigs, caused by an operation performed on the spinal chord, 1869

of epilepsy produced by injuring the spinal chord of guinea-pigs, being inherited, 1861 1866

in guinea-pigs, of the inherited effects of operations, 1872

→ cautious in denying such power. 1866 1869

cautious. 1861

cautious in denying this tendency. 1872

→ Hence it will perhaps be safest to look at 1869 1872

There is not sufficient evidence to induce us to believe that mutilations are ever inherited; and I should prefer explaining 1859

There is not sufficient evidence to induce me to believe that mutilations are ever inherited; and I should prefer explaining 1860

So that it will perhaps be safest to look at 1861 1866

→ as due to the 1869

by the long-continued 1859 1860

as due to the long-continued 1861 1866

not as cases of inherited mutilations, but as due to the 1872

→ long-continued disuse; 1869 1872

disuse in their progenitors; 1859 1860 1861 1866

→ OMIT 1869 1872

the tarsi are almost always lost in 1859 1860 1861 1866

→ are generally found with their tarsi lost, this must happen 1869 1872

they must be lost 1859 1860 1861 1866

→ the tarsi cannot be of much importance or 1869 1872

cannot 1859 1860 1861 1866

→ much 1859 1860 1869 1872

of much importance or be much 1861 1866

which

have

lately inhabited several oceanic islands, tenanted by no beast of prey, has been caused by disuse. The ostrich indeed inhabits

continents,

continents

and is exposed to danger from which it cannot escape by flight, but

~~by kicking~~

by kicking

it can defend itself →from enemies, as

efficiently

well

many

any

→of the smaller quadrupeds. We may

imagine

believe

that the

early

....

progenitor of the ostrich

had

genus had

habits like those of

the

bustard, and

that

that,

as →natural selection increased in successive generations the size and weight of its

body

body,

→its legs were used more, and its wings less, until they became incapable of flight.

Kirby has remarked (and I have observed the same fact) that the anterior tarsi, or feet, of many male dung-feeding beetles are

~~very~~

very

often broken off; he examined seventeen specimens in his own collection, and not one had even a relic left. In the Onites apelles the tarsi are so habitually lost, that the insect has been described as not having them. In some other genera they are present, but in a rudimentary condition. In the Ateuchus or sacred

bettle

beetle

of the Egyptians, they are totally deficient. The evidence that accidental mutilations can be inherited is at present →not quite decisive; but the remarkable

cases

case

observed by Brown-Séquard →of inherited epilepsy in guinea-pigs, caused by an operation performed on the spinal chord, should make us →cautious in denying such power. →Hence it will perhaps be safest to look at the entire absence of the anterior tarsi in Ateuchus, and their rudimentary condition in some other genera, →as due to the effects of →long-continued disuse; for as →OMIT many dung-feeding

beetles,

beetles

→are generally found with their tarsi lost, this must happen early in

life,

life;

and

....

therefore →the tarsi cannot be of much importance or be →much used by these insects.

In some cases we might easily put down to disuse modifications of structure which are wholly, or mainly,