Origin of Species Variorum

→ formerly within each isolated 1869

in each broken 1859 1860 1861

within each isolated 1866 1872

→ during the process of natural selection will have been supplanted and exterminated, 1866 1869 1872

will have been supplanted and exterminated during the process of natural selection, 1859 1860 1861

→ together all the species of the same group, 1866 1869 1872

all the species of the same group together, 1859 1860 1861

→ in a future chapter attempt to show, in 1859 1860 1861 1866 1869

attempt to show in a future chapter, in 1872

separately been rendered sufficiently distinct to rank as representative species. In this case, intermediate varieties between the several representative species and their common parent, must

formerly

....

have existed →formerly within each isolated portion of the land, but these links →during the process of natural selection will have been supplanted and exterminated, so that they will no longer

exist

be found

in a living state.

Thirdly, when two or more varieties have been formed in different portions of a strictly continuous area, intermediate varieties will, it is probable, at first have been formed in the intermediate zones, but they will generally have had a short duration. For these intermediate varieties will, from reasons already assigned (namely from what we know of the actual distribution of closely allied or representative species, and likewise of acknowledged varieties), exist in the intermediate zones in lesser numbers than the varieties which they tend to connect. From this cause alone the

interme- diate

intermediate

varieties will be liable to accidental extermination; and during the process of further modification through natural selection, they will almost certainly be beaten and supplanted by the forms which they connect; for these from existing in greater numbers will, in the aggregate, present more

varieties,

variation,

and thus be further improved through natural selection and gain further advantages.

Lastly, looking not to any one time, but to all time, if my theory be true, numberless intermediate varieties, linking

most

....

closely →together all the species of the same group, must assuredly have existed; but the very process of natural selection constantly tends, as has been so often remarked, to exterminate the

parent forms

parent-forms

and the intermediate links. Consequently evidence of their former existence could be found only amongst fossil remains, which are preserved, as we shall →in a future chapter attempt to show, in