Origin of Species Variorum

→ ants or parents, 1869 1872

parents, 1859 1860 1861 1866

→ jaws having a 1859 1860 1861 1866 1869

OMIT 1872

→ at the same time 1869

simultaneously 1859 1860 1861 1866 1872

→ OMIT 1866 1869 1872

from being the most useful to the community, 1859 1860 1861

→ in greater and greater numbers, 1869 1872

in greater and greater numbers 1859 1860 1861

OMIT 1866

→ until 1859 1860 1861 1869 1872

in greater and greater numbers, until 1866

→ OMIT 1869 1872

at the same time and place 1866

→ I have now explained how, 1866 1869 1872

Thus, 1859 1860 1861

the working ants of the several sizes differed wonderfully in shape, and in the form and number of the teeth. But the important fact for us is,

that

that,

though the workers can be grouped into castes of different sizes, yet they graduate insensibly into each other, as does the widely-different structure of their jaws. I speak confidently on this latter point, as

Mr.

Sir J.

Lubbock made drawings for

me,

with the camera

lucida

lucida,

of the jaws which I

had

....

dissected from the workers of the several sizes. Mr. Bates, in his

~~most~~

most

interesting 'Naturalist on the Amazons,' has described

some

....

analogous cases.

With these facts before me, I believe that natural selection, by acting on the fertile →ants or parents, could form a species which should regularly produce neuters,

~~either~~

either

all of large size with one form of jaw, or all of small size with →jaws having a widely different

jaws;

structure;

or lastly, and this is

our

the

climax of difficulty, one set of workers of one size and structure, and →at the same time another set of workers of a different size and structure;— a graduated series having

been first

first been

formed, as in the case of the driver ant, and then the extreme

forms,

forms

→OMIT having been produced →in greater and greater numbers, through the

natural selection

survival

of the parents which generated

them;

them,

→until none with an intermediate structure were produced.

An analogous explanation has been given by Mr. Wallace, of the equally complex case, of certain Malayan

butterflies

Butterflies

regularly appearing →OMIT under two or even three distinct female forms; and by Fritz

Müller,

of certain Brazilian crustaceans likewise appearing under two widely distinct male forms. But

the

this

subject need not here be discussed.

→I have now explained how, as I believe, the wonderful fact of two distinctly defined castes of sterile workers existing in the same nest, both widely different from each other and from