Origin of Species Variorum

→ just be distinguished, yet the 1869

be distinguished from each other, yet their 1859 1860 1861 1866

just be distinguished, the 1872

→ in all its proportions from 1869

from 1859 1860 1861 1866 1872

→ OMIT 1869

in all its proportions, 1859 1860 1861 1866

in 1872

→ as much 1859 1860 1861 1866 1869

the same proportions 1872

→ The 1859 1860 1861 1866 1869

These facts are explained by the above 1872

→ above two principles 1869

two principles above given seem to me to 1859 1860 1861

two principles above given, namely that variations do not generally supervene at a very early age, and that they are inherited at a corresponding age whatever that may have been, seem to me to 1866

two principles. 1872

OMIT 1872

facts in regard to the later embryonic stages of our domestic varieties. Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been acquired earlier or later in life, if the full-grown animal possesses them. 1859 1860 1861

several facts regarding the later developmental stages of our domestic varieties. 1866

→ And the cases just given, more especially that of pigeons, show that the characteristic differences which give value to the breeds and which have been 1869

And the cases just given, more especially that of pigeons, seem to show that the characteristic differences which give value to each breed, and which have been 1859 1860 1861

Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been 1866

Fanciers select their dogs, horses, pigeons, &c., for breeding, when nearly grown up: they are indifferent whether the desired qualities are 1872

→ accumulated by mans selection, have not generally appeared at a very early period of life, and have been inherited at a corresponding not early period. 1869

accumulated by man's selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period. 1859 1861

accumulated by mans selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period. 1860

acquired earlier or later in life, if the full-grown animal possesses them. 1866 1872

→ of the short-faced tumbler, which when twelve hours old possessed its proper characters, proves that this is not the universal rule; for here 1869

of the short-faced tumbler, which when twelve hours old had acquired its proper proportions, proves that this is not the universal rule; for here 1859 1860 1861

just given, more especially that of pigeons, seem to show that 1866

just given, more especially that of the pigeons, show that 1872

→ must either have appeared at an earlier period than usual, or, if not so, the differences must 1859 1860 1861 1869

which give value to each breed, and which 1866

which 1872

→ inherited, not at a corresponding, but 1869

inherited, not at the corresponding, but 1859 1860 1861

accumulated by man's selection, have not generally first appeared 1866

accumulated by man's selection, and which give value to his breeds, do not generally appear 1872

→ earlier age. 1859 1860 1861 1869

early period of life, and have been inherited by the offspring at a corresponding not early period. 1866

early period of life, and are inherited at a corresponding not early period. 1872

↑ 1 blocks not present in 1859 1860 1861 1869; present in 1866 1872

But the case of the short-faced tumbler, which when twelve hours old had acquired its proper proportions, proves that this is not the universal rule; for here the characteristic differences must either have appeared at an earlier period than usual, or, if not so, the differences must have been inherited, not at the corresponding, but at an earlier age.

→ two principles 1869 1872

facts and the above two principles— which latter, though not proved true, can be shown to be in some degree probable— 1859 1860 1861

facts and the above two principles 1866

→ OMIT 1869 1872

on my theory 1859 1860 1861 1866

→ ancient form and 1869 1872

one parent-species, and of which the several new species have become 1859 1860 1861 1866

→ for different 1869 1872

in accordance with their diverse 1859 1860 1861 1866

→ in the several species at a not early 1869 1872

at a rather late 1859 1860 1861 1866

→ will be left but little modified and will 1869

of the new species of our supposed genus will manifestly tend to 1859 1860 1861 1866

will have been but little modified, and they will still 1872

the nestling birds of these several breeds were placed in a row, though most of them could →just be distinguished, yet the proportional differences in the above specified

several

....

points were incomparably less than in the full-grown birds. Some characteristic points of difference— for instance, that of the width of mouth— could hardly be detected in the young. But there was one remarkable exception to this rule, for the young of the short-faced tumbler differed →in all its proportions from the young of the wild rock-pigeon and of the other breeds, →OMIT almost exactly →as much as in the adult state.

→The →above two principles

~~explain~~

explain

~~these~~

these

→facts. Fanciers select for breeding their dogs, horses, pigeons, &c., when nearly grown up: they are indifferent whether the desired qualities are acquired earlier or later in life, if the full-grown animal possesses them. →And the cases just given, more especially that of pigeons, show that the characteristic differences which give value to the breeds and which have been →accumulated by mans selection, have not generally appeared at a very early period of life, and have been inherited at a corresponding not early period.

And

But

the

cases

case

→of the short-faced tumbler, which when twelve hours old possessed its proper characters, proves that this is not the universal rule; for here the characteristic differences →must either have appeared at an earlier period than usual, or, if not so, the differences must have been →inherited, not at a corresponding, but at

a very

→earlier age. ↑

Now let us apply these →two principles to species in a state of nature. Let us take a

genus

group

of birds, descended →OMIT from some →ancient form and modified through natural selection →for different habits. Then, from the many slight successive

steps of

....

variation

variations

having supervened →in the several species at a not early age, and having been inherited at a corresponding age, the young →will be left but little modified and will