| → we doubt that the variations or individual differences, which are 1869 | 
| we doubt that variations 1859 1860 1861 1866 | 
| not variations or individual differences, which are 1872 | 
  | 
| → beneficial would 1869 | 
| useful to beings, under their excessively complex relations of life, would 1859 1860 1861 | 
| useful to them under their extremely complex relations of life would 1866 | 
| beneficial, 1872 | 
  | 
| → be preserved and accumulated through natural selection, or the survival of the fittest? 1869 1872 | 
| be preserved, accumulated, and inherited? 1859 1860 1861 | 
| occasionally occur, and then be preserved and accumulated by natural selection? 1866 | 
  | 
| → him, why, 1869 1872 | 
| himself, should nature fail in selecting variations useful, 1859 1860 1861 | 
| himself, should nature fail in preserving or selecting variations useful, 1866 | 
  | 
| → and complex conditions 1869 1872 | 
| conditions 1859 1860 1861 1866 | 
  | 
| → should not variations useful to natures 1869 | 
| to her 1859 1860 1861 1866 | 
| should not variations useful to nature's 1872 | 
  | 
| → products often arise, and be preserved or selected? 1869 1872 | 
| products? 1859 1860 1861 1866 | 
  | 
| → itself 1859 1860 1861 1866 1869 | 
| the highest degree 1872 | 
  |