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It has been disputed at what period of life the causes of variability, whatever they may be, generally act; whether during the early or late period of development of the embryo, or at the instant of conception.

the same manner. Such facts as the complex and extraordinary out-growths which invariably follow from the insertion of a minute drop of poison by a gall-producing insect, show us what singular modifications might result in the case of plants from a chemical change in the nature of the sap.
Indefinite variability is a much more common result of changed conditions than definite variability, and has probably played a more important part in the formation of our domestic races. We see indefinite variability in the endless slight peculiarities which distinguish the individuals of the same species, and which cannot be accounted for by inheritance from either parent or from some more remote ancestor. Even strongly-marked differences occasionally appear in the young of the same litter, and in seedlings from the same
seed-capsule.
seed-capsules.
At long intervals of time, out of millions of individuals reared in the same country and fed on nearly the same food, deviations of structure so strongly pronounced as to deserve to be called monstrosities arise; but monstrosities cannot be separated by any distinct line from slighter variations. All such changes of structure, whether extremely slight or strongly marked, which appear amongst many individuals living together, may be considered as the indefinite effects of the conditions of life on each individual organism, in nearly the same manner as a chill affects different men in an indefinite manner, according to their state of body or constitution, causing coughs or colds, rheumatism, or inflammations of various organs.
With respect to what I have called the
in direct
indirect
action of changed conditions, namely, through the reproductive system being affected, we may infer that variability is thus induced, partly from the fact of this system being extremely sensitive to any change in the conditions,