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labour and wax 1869 1872
wax 1859 1860 1861 1866

would 1859 1860 1861 1869 1872
and labour would 1866

labour and wax. 1866 1869 1872
wax. 1859 1860 1861

plates; 1866 1869 1872
plates. 1859 1860
plates. The motive power of the process of natural selection having been economy of wax, together with cells of due strength, and of the proper size and shape for the larvæ; that individual swarm which made the best cells, and wasted least honey in the secretion of wax, having succeeded best, and having transmitted by inheritance their newly acquired economical instincts to new swarms, which in their turn will have had the best chance of succeeding in the struggle for existence. 1861

the construction of cells of due strength and of the proper size and shape for the larvæ, this being effected with the greatest possible economy of labour and wax; 1869 1872
economy of wax; 1859 1860
the construction of cells of due strength and of the proper size and shape for the larvæ, this being effected with the greatest possible economy of was and labour; 1866

thus made the best cells with least labour, and least waste of 1866 1869 1872
wasted least 1859 1860

their 1872
by inheritance its 1859 1860
by inheritance their 1866 1869

to our
humble-bee,
humble-bees,
if
she
they
were to make
her
their
cells more and more regular, nearer together, and aggregated into a mass, like the cells of the Melipona; for in this case a large part of the bounding surface of each cell would serve to bound
other
the adjoining
cells, and much labour and wax would be saved. Again, from the same cause, it would be advantageous to the Melipona, if she were to make her cells closer together, and more regular in every way than at present; for then, as we have seen, the spherical surfaces would wholly
disappear,
disappear
and
would all
....
be replaced by plane surfaces; and the Melipona would make a comb as perfect as that of the hive-bee. Beyond this stage of perfection in architecture, natural selection could not lead; for the comb of the hive-bee, as far as we can see, is absolutely perfect in economising labour and wax.
Thus, as I believe, the most wonderful of all known instincts, that of the hive-bee, can be explained by natural selection having taken advantage of numerous, successive, slight modifications of simpler instincts; natural selection
having
having,
by slow degrees, more and more
perfectly,
perfectly
led the bees to sweep equal spheres at a given distance from each other in a double layer, and to build up and excavate the wax along the planes of
intersection.
intersection;
The
the
bees, of course, no more knowing that they swept their spheres at one particular distance from each other, than they know what are the several angles of the hexagonal prisms and of the basal rhombic plates;
The
the
motive power of the process of natural selection having been the construction of cells of due strength and of the proper size and shape for the larvæ, this being effected with the greatest possible economy of labour and wax; that individual swarm which thus made the best cells with least labour, and least waste of honey in the secretion of wax, having succeeded best, and having transmitted their
newly acquired
newly-acquired
economical
instinct
instincts
to new swarms, which in their turn will have had the best chance of succeeding in the struggle for existence.
Objections to the Theory of Natural Selection as applied to Instincts: Neuter and Sterile
Insects .
Insects .
It has been objected to the foregoing view
on
of
the origin of
instinct
instincts
that "the variations of structure and of instinct must have been simultaneous and accurately adjusted to each other, as a modification in the one without an immediate corresponding change in the other would have been fatal." The force of this objection
seems
rests
entirely
to rest
....
on the assumption that the changes in
both
the
instinct
instincts
and structure are abrupt. To take as an illustration the case of the larger titmouse (Parus major) alluded to in
the last
a previous
chapter: