See page in:
1859
1860
1861
1866
1869
1872

Compare with:
1859
1860
1861
1866
1869

applied to the stigma of some one species of the same genus, 1872
of the same genus applied to the stigma of some one species, 1859 1860 1861 1866
of the same genus applied to the stigma of some one of the species, 1869

The hybrids raised 1872
Hybrids 1859 1860 1861 1866 1869

as in the genus Verbascum, can 1869 1872
can 1859 1860 1861 1866

in degree when 1872
when 1859 1860 1861 1866 1869

fertility, the pollen of
differ- ent
different
species applied to the stigma of some one species of the same genus, yields a perfect gradation in the number of seeds produced, up to nearly complete or even quite complete fertility; and, as we have seen, in certain abnormal cases, even to an excess of fertility, beyond that which the
plants
plant's
own pollen
will
....
produce.
produces.
So in hybrids themselves, there are some which never have produced, and probably never would produce, even with the pollen of
either
the
pure
parent,
parents,
a single fertile seed: but in some of these cases a first trace of fertility may be detected, by the pollen of one of the pure parent-species causing the flower of the hybrid to wither earlier than it otherwise would have done; and the early withering of the flower is well known to be a sign of incipient
fertilization.
fertilisation.
From this extreme degree of sterility we have self-fertilised hybrids producing a greater and greater number of seeds up to perfect fertility.
The hybrids raised from two species which are very difficult to cross, and which rarely produce any offspring, are generally very sterile; but the parallelism between the difficulty of making a first cross, and the sterility of the hybrids thus
produced—two
produced—
two
two
classes of facts which are generally confounded
together—is
together—
is
is
by no means strict. There are many cases, in which two pure
species
species,
as in the genus Verbascum, can be united with unusual facility, and produce numerous hybrid-offspring, yet these hybrids are remarkably sterile. On the other hand, there are species which can be crossed very rarely, or with extreme difficulty, but the hybrids, when at last produced, are very fertile. Even within the limits of the same genus, for instance in Dianthus, these two opposite cases occur.
The fertility, both of first crosses and of hybrids, is more easily affected by unfavourable conditions, than is
the fertility
that
of pure species. But the
degree
fertility
of
fertility
fer- tility
first crosses
is likewise innately variable; for it is not always the same in degree when the same two species are crossed under the same
circumstances,
circumstances;
but
it
depends in part upon the constitution of the individuals which happen to have been chosen for the experiment. So it is with hybrids, for their degree of fertility is often found to differ greatly in the several individuals raised from seed out of the same capsule and exposed to
exactly
....
the same conditions.
By the term systematic affinity is meant, the
resemblance
general resemblance
between species in structure and
in
....
constitution,
constitution.
more
....
especially
....
in
....
the
....
structure
....
of
....
parts
....
which
....
are
....
of
....
high
....
physiological
....
importance
....
and
....
which
....
differ
....
little
....
in
....
the
....
allied
....
species.
....
Now the fertility of first
crosses
crosses,
between species,
....
and of the hybrids produced from them, is largely governed by their systematic affinity. This is clearly shown by hybrids never having been raised between species ranked by systematists in distinct families; and on the other hand, by very