Origin of Species Variorum

→ namely, that slight variations generally appear at a not very early period of life, and are inherited at a corresponding not early period, explain, as 1869 1872

if their truth be admitted, will, 1859 1860 1861 1866

→ varieties, 1869 1872

and have probably 1859 1860 1861 1866

→ race-horses— breeds which have been almost wholly formed by selection under domestication— 1869 1872

race-horses 1859 1860 1861 1866

→ OMIT 1869 1872

and this surprised me greatly, as I think it probable that the difference between these two breeds has been wholly caused by selection under domestication; 1859 1860 1861 1866

→ this is 1869 1872

the colts have 1859 1860 1861 1866

→ the case. 1869 1872

acquired their full amount of proportional difference. 1859 1860 1861 1866

→ we have conclusive evidence 1869 1872

the evidence appears to me conclusive, 1859 1860 1861 1866

→ the Pigeon are 1869 1872

Pigeon have 1859 1860 1861 1866

→ the young 1869 1872

young pigeons of various breeds, 1859 1860 1861 1866

→ the 1859 1860 1861 1866 1872

in the wild parent-species, in pouters, fantails, runts, barbs, dragons, carriers, and tumblers, the 1869

→ in so extraordinary a manner in the 1872

so extraordinarily in 1859 1860 1861 1866

in so extraordinary a degree in the 1869

→ and in other characters, that 1869 1872

that 1859 1860 1861 1866

→ certainly have been ranked as 1866 1872

I cannot doubt, be ranked in 1859 1860 1861

certainly be ranked as 1869

→ if found in a state of nature. 1869 1872

had they been natural productions. 1859 1860 1861 1866

→ just be distinguished, the 1872

be distinguished from each other, yet their 1859 1860 1861 1866

just be distinguished, yet the 1869

→ from 1859 1860 1861 1866 1872

in all its proportions from 1869

→ in 1872

in all its proportions, 1859 1860 1861 1866

OMIT 1869

→ the same proportions 1872

as much 1859 1860 1861 1866 1869

→ These facts are explained by the above 1872

The 1859 1860 1861 1866 1869

→ two principles. 1872

two principles above given seem to me to 1859 1860 1861

two principles above given, namely that variations do not generally supervene at a very early age, and that they are inherited at a corresponding age whatever that may have been, seem to me to 1866

above two principles 1869

→ OMIT 1872

facts in regard to the later embryonic stages of our domestic varieties. Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been acquired earlier or later in life, if the full-grown animal possesses them. 1859 1860 1861

several facts regarding the later developmental stages of our domestic varieties. 1866

facts. Fanciers select for breeding their dogs, horses, pigeons, &c., when nearly grown up: they are indifferent whether the desired qualities are acquired earlier or later in life, if the full-grown animal possesses them. 1869

→ Fanciers select their dogs, horses, pigeons, &c., for breeding, when nearly grown up: they are indifferent whether the desired qualities are 1872

And the cases just given, more especially that of pigeons, seem to show that the characteristic differences which give value to each breed, and which have been 1859 1860 1861

Fanciers select their horses, dogs, and pigeons, for breeding, when they are nearly grown up: they are indifferent whether the desired qualities and structures have been 1866

And the cases just given, more especially that of pigeons, show that the characteristic differences which give value to the breeds and which have been 1869

→ acquired earlier or later in life, if the full-grown animal possesses them. 1866 1872

accumulated by man's selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period. 1859 1861

accumulated by mans selection, have not generally first appeared at an early period of life, and have been inherited by the offspring at a corresponding not early period. 1860

accumulated by mans selection, have not generally appeared at a very early period of life, and have been inherited at a corresponding not early period. 1869

→ just given, more especially that of the pigeons, show that 1872

of the short-faced tumbler, which when twelve hours old had acquired its proper proportions, proves that this is not the universal rule; for here 1859 1860 1861

just given, more especially that of pigeons, seem to show that 1866

of the short-faced tumbler, which when twelve hours old possessed its proper characters, proves that this is not the universal rule; for here 1869

These two principles, →namely, that slight variations generally appear at a not very early period of life, and are inherited at a corresponding not early period, explain, as I believe,

explain

....

all the above specified leading facts in embryology. But first let us look

a few analogous cases in

domestic

our domestic

varieties. Some authors who have written on Dogs, maintain that the greyhound and

bull-dog,

bulldog,

though

appearing

....

so different, are really

varieties most

varieties

....

closely

allied,

allied

→varieties, descended from the same wild stock; hence I was curious to see how far their puppies differed from each other: I was told by

breeder

breeders

that they differed just as much as their parents, and this, judging by the eye, seemed almost to be the case; but on actually measuring the old dogs and their

six-days old

six-days-old

puppies, I found that the puppies had not

nearly acquired

acquired nearly

their full amount of proportional difference. So, again, I was told that the foals of cart and →race-horses— breeds which have been almost wholly formed by selection under domestication— differed as much as the full-grown animals; →OMIT but having had careful measurements made of the

dam

dams

and of

~~the~~

three-days

old

colt

a race

race-horses

race

and heavy

cart-horse,

cart-horses,

I find that →this is by no means →the case.

As →we have conclusive evidence that the

several domestic

....

breeds of →the Pigeon are descended from

one

a single

wild species, I compared →the young within twelve hours after being hatched; I carefully measured →the proportions (but will not here give

details)

the details)

of the beak, width of mouth, length of nostril and of eyelid, size of feet and length of

leg.

leg,

~~the~~

the

~~wild~~

wild

stock,

~~parent-species,~~

~~pouters,~~

~~fantails,~~

~~runts,~~

~~barbs,~~

~~dragons,~~

~~carriers,~~

~~and~~

~~tumblers.~~

Now some of these birds, when mature, differ →in so extraordinary a manner in the length and form of beak, →and in other characters, that they

would,

would

→certainly have been ranked as distinct

genera,

genera

→if found in a state of nature. But when the nestling birds of these several breeds were placed in a row, though most of them could →just be distinguished, the proportional differences in the above specified

several

....

points were incomparably less than in the full-grown birds. Some characteristic points of difference— for instance, that of the width of mouth— could hardly be detected in the young. But there was one remarkable exception to this rule, for the young of the short-faced tumbler differed →from the young of the wild rock-pigeon and of the other breeds, →in almost exactly →the same proportions as in the adult state.

→These facts are explained by the above →two principles.

explain

....

these

....

→OMIT →Fanciers select their dogs, horses, pigeons, &c., for breeding, when nearly grown up: they are indifferent whether the desired qualities are →acquired earlier or later in life, if the full-grown animal possesses them.

But

And

the

case

cases

→just given, more especially that of the pigeons, show that