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1859
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1869
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1859
1860
1861
1866
1869

doubtful or at last into well-defined 1872
well-defined 1859 1860 1861 1866
doubtful or at last well-defined 1869

m 14 , 1859 1861 1866 1869 1872
m 14 1860

z 10 ) 1859 1861 1866 1869 1872
z 10 1860

z 14 , 1859 1861 1866 1869 1872
z 14 1860

In any genus, the species 1869 1872
In each genus, the species, 1859 1860
The species of a genus 1861 1866

from each other, will 1869 1872
will 1859 1860 1861 1866

but unequal periods continue to transmit 1869 1872
period continue transmitting 1859
period continue to transmit 1860 1861 1866

more numerous or greater in amount, to convert these three forms into doubtful or at last into well-defined
species:
species.
thus
Thus
the diagram illustrates the steps by which the small differences distinguishing varieties are increased into the larger differences distinguishing species. By continuing the same process for a greater number of generations (as shown in the diagram in a condensed and simplified manner), we get eight species, marked by the letters between a 14 and m 14 ,
, all
all
descended from (A). Thus, as I believe, species are multiplied and genera are formed.
In a large genus it is probable that more than one species would vary. In the diagram I have assumed that a second species (I) has produced, by analogous steps, after ten thousand generations, either two well-marked varieties ( w 10 and z 10 )
) or
or
two species, according to the amount of change supposed to be represented
be-
....
tween
between
the horizontal lines. After fourteen thousand generations, six new species, marked by the letters n 14 to z 14 ,
, are
are
supposed to have been produced. In any genus, the species which are already
extremely
very
different in
character,
character
from each other, will generally tend to produce the greatest number of modified descendants; for
they
these
will have the best chance of
filling
seizing on
new and widely different places in the polity of nature: hence in the diagram I have chosen the extreme species (A), and the nearly extreme species (I), as those which have largely varied, and have given rise to new varieties and species. The other nine species (marked by capital letters) of our original genus, may for
a
....
long but unequal periods continue to transmit unaltered
descendants:
descendants;
and this is shown in the diagram by the dotted lines
not
unequally
prolonged
far
....
upwards
upwards.
from
....
want
....
of
....
space.
....
But during the process of modification, represented in the diagram, another of our principles, namely that of extinction, will have played an important part. As in each fully stocked country natural selection necessarily acts by the selected form having some advantage in the struggle for life over other forms, there will be a constant tendency in the improved descendants of any one species to supplant and exterminate in each stage of descent their predecessors and their original
parent.
progenitor.
For it should be remembered that the competition will generally be most severe between those forms which are most nearly related to each other in habits, constitution, and structure. Hence all the intermediate forms between the earlier and later states, that is between the less and more improved
state
states
of
a
the same
species, as well as the original parent-species itself, will generally tend to become extinct. So it probably will be with many whole collateral lines of descent, which will be conquered by later and improved
lines
lines.
of
....
descent.
....
If, however, the modified offspring of a species get into some distinct country, or become quickly adapted to some