Reproducing images from Darwin Online

Some of the publications with illustrations from Darwin Online

In order to reproduce materials from Darwin Online in another publication or broadcast, written permission must be sought from the Director, Dr John van Wyhe. Specify exactly which image(s) or transcription you would like to reproduce (with the URL, e.g. or submit a screenshot.

Frequently reproduced images can be found here.

See also: The Complete Photographs of Darwin.

Permission is not required for referenced quotations, use in talks and presentations or unpublished exhibition posters, student papers and theses or to link to Darwin Online from another website. High-resolution images are available for many images for a one-time 50GBP administration charge per image.

Note: Darwin Online cannot give permission to reproduce most of the manuscripts, that is, hand-written documents, on the site. Such requests must be sent to the owner of the manuscript. For most manuscripts see the Cambridge University Library Order Form for Digital Images (such images are prefaced here with CUL-). Certain other materials, for which Darwin Online has received reproduction permission, may not be reproduced without the permission of the original copyright holder. Therefore it is necessary to request permission and specify the image(s).

Works containing images from Darwin Online include:

Films, documentaries, tv programmes, books, articles, stamps, etc.

Limited Edition Casino quality twin-deck Playing Cards commemorating Darwin's Bicentenary and 150 years of The Origin of Species. Illustrated with images from Darwin Online (click here for a sample).

Ten new Darwin stamps. The Royal Mail. 12 Feb. 2009. − a Galapogos-themed miniature sheet of four stamps pays homage to the influence of the islands on Darwin's thought, while a set of six 'jigsaw' design stamps explore the breadth of the influence of Darwin's idea's across the natural sciences. The two sets of stamps will be issued on 12 February, with the first day issues also having special post marks designed for the occasion.

The Portuguese Post Office issued a set of seven Darwin bicentenary stamps featuring images from Darwin Online.

Darwin 200th Anniversary issue. The Gibraltar Philatelic Bureau issued five stamps using images from Darwin Online. click here

Cocos (Keeling) Islands stamps, 55c - 1836 Charles Darwin visited the islands on The Beagle, using images from Darwin Online.

Limited Edition Casino quality twin-deck Playing Cards commemorating Darwin's Bicentenary and 150 years of The Origin of Species. Illustrated with images from Darwin Online (click here for a sample).

The Royal Mail. Ten new Darwin stamps. 12 Feb. 2009. − a Galapogos-themed miniature sheet of four stamps pays homage to the influence of the islands on Darwin's thought, while a set of six 'jigsaw' design stamps explore the breadth of the influence of Darwin's idea's across the natural sciences. The two sets of stamps will be issued on 12 February, with the first day issues also having special post marks designed for the occasion.

The Portuguese Post Office issued a set of seven Darwin bicentenary stamps featuring images from Darwin Online.

The Gibraltar Philatelic Bureau Darwin 200th Anniversary issue: five stamps using images from Darwin Online. click here

Cocos (Keeling) Islands stamps, 55c - 1836 Charles Darwin visited the islands on The Beagle, using images from Darwin Online.

"Great Nature" on NHK (Japanese public TV) 2015.

3sat nano (German public television), 11.2018.

Aharoni, Prof. Ron. Why do we frown? (Hebrew).

Ambasciano, Leonardo. History as a Canceled Problem?

American Association for the Advancement of Science. 2009. Darwin 200 collection. Science Magazine 2009.

Andalucia Innova (special issue on human evolution) [Spain]

Anon. 2009. Darwin bicentenary celebration. Galapagos Conservation Trust, UK News, Spring/Summer 2009, p. 4.

Anon. 2009. Darwin bicentenary celebration. Galapagos Conservation Trust, UK News, Spring/Summer 2009, p. 4.

Anon. 2009. Samla pa Darwiniana. Penséer (Stockholm), no. 2: 4-13.

Arsuaga, Juan Luis. 2008. Darwin, el naturalista. Accentura, pp. 1-174.

Atheism (documentary, Juniper productions) 2006.

Attenborough, David. 2009. Charles Darwin and the Tree of Life, BBC One. one part tv documentary. 2009, 02.01.

Attenborough, David. 2009. Life Stories.

Attenborough, David. 2013. Plants behaving badly. 2 episodes. Produced by Sky Vision for Terra Mater Factual Studios. (BskyB).

Attenborough, David. 2018. Life on Earth. New edn.

Attenborough, David. 2018. The greatest story ever told: Life on Earth. William Collins. Hardback.

Ault, Kip. 2016. Do Elephants Have Knees? Serious Whimsy in Darwinian Stories of Origins. Cornell University Press.

Ault, Kip. 2021. On Beyond Science Standards.

Azania – archaeological research in Africa, 2020.

Baker, Abigail. Discovering Troy. Bloomsbury Academic.

Bartolomé, Gerardo & Glickman, Barry. 2008. Patagonia through the eyes of Darwin, [in English and Spanish], Zagier & Urruty Publications.

Baur, Bruno. 2010. Biodiversität. Berlin, Stuttgart, Wien: Haupt Verlag.

Baur, Bruno. text book on Conservation Biology. Haupt Verlag, Switzerland.

Baxter, Craig, Juan v. Martinez Luciano, 2009. Fiebre y Credo: Correspondencia entre Darwin y Gray.

BBC Earth. Online film commissioned. 2021

BBC Horizon – 'Oceans of the Solar System' [Mars], 2015.

BBC4 production – A Garden in Time. 

Bear, Mark F., Barry W. Connors, and Michael A. Paradiso. 2014. Neuroscience Exploring the Brain, Fourth Edition. Lippincott Williams & Wilkins.

Beekman, Madeleine. How Evolution Works.

Berkeley Scientific Journal, Fall 2020.

Binney, A. J. and Andrew Plant. 2021. Mr Darwin's Birds.

Blair, 2017. "Settler Indigeneity and the Eradication of the Non-Native: Self-Determination and Biosecurity in the Falkland Islands (Malvinas)." The Journal of the Royal Anthropological Institute (JRAI). vol. 23 (3).

Bombaro, C. 2012. Finding history: research methods and resources for students and scholars. Scarecrow press.

Books that changed History. (Dorling Kindersley 2017) [cover image, spine of Origin 1859]

Bouzat, Juan Luis. 2014. Darwin's Diagram of Divergence of Taxa as a Causal Model for the Origin of Species. The Quarterly Review of Biology (March) 89, no. 1: 21-38. (without acknowledgement)

Bradley, J. T. and J. Lamar. 2013. Charles Darwin a celebration of his life and legacy.

Brown et al. 2014. First glacier inventory and recent glacier variations on Isla Grande de Tierra del Fuego and adjacent islands in Southern Chile. Springer.

Browne, Janet. 2018. The quotable Darwin. Princeton University Press.

Bruggs, Richard. 2017, 05. Darwin's abominable mystery. Nature ecology & evolution.

Busk, Meningen med Livet: Om katusfinker, evolutions og kaos. 2013.

Bustamante, Luis Arana. 2016. Sobre la estadía de Darwin en el Perú (1835) Un avance de investigación. Investigaciones sociales. Vol. 20 N°36, pp.151-157.

Bustamante, Luis Arana. 2017. Conociendo a Charles Darwin (1809 - 1882) Folia Darwiniana 1: 9-22.

Bustamante, Luis Arana. 2017. I extracted these remains by digging a hole...: Charles Darwin, protoarqueólogo en Lima [1835]. Un avance de investigación. Boletín YUNGAS 1(3). September: 3-7.

Bustamante, Luis Arana. 2018. Darwin en el Perú (1835): un panorama de los primeros días. Folia Darwiniana 3:27-38.

Cam. No. 56, Lent Term, 2009.

Carroll, Sean. 2018. The Story of Life. W.W. Norton.

Carroll, Sean. 2020. By Chance: Asteroids, Genes, and Humanity.

Cartwright, 2016. Evolution and Human behaviour, Palgrave.

Carver, Alternate Histories and Nineteenth-Century Literature: Untimely Meditations in Britain, France and America. 2017.

Catalyst, seconday science review, vol. 19 no. 4, April 2009. With papers by Katie Edwards (Celebrating Darwin), Gary Skinner (A box of beetles), John Stacey (The Socotra Archipelago), Gary Skinner (The big picture: Charles Darwin at Down House), Peter Bentley (Darwin's digital children), and a page on the Beagle Voyage.

Cavin, Lionel. 2009. Darwin et les fossiles: histoire d'une réconciliation. Chene-Bourg: Georg Editeur, pp. 1-237.

CBC Radio-Canada. TV documentary on Darwin. (French). 2009.

Chancellor, Gordon and John van Wyhe eds. with the assistance of Kees Rookmaaker. 2009. Charles Darwin's notebooks from the voyage of the Beagle. With a foreword by Richard Darwin Keynes. Cambridge University Press.

Charles Darwin and evolution. 1809-2009. Website by students at Christ's College, Cambridge.

Charles Darwin University. Origins 2009 edition 2. (September).

Charles Darwin. Environnement. (XD Productions, France). 2009. (documentary)

CHIKO-CHAN, NHK (Japan TV). 2018.

Cite-sciences. 2016. Accompagnement de visite de l'exposion Darwin, l'original.

Clement, J. 2009. Darwin's notebook: the life, times, and discoveries of Charles Robert Darwin.

Clifford B. Frith, 2016. Charles Darwin's Life With Birds. His Complete Ornithology. OUP.

Cohen, Claudine. 2010. Darwin on woman. Darwin et la femme. Comptes Rendus Biologies 333, Issue 2 (February): 157-165.*

Complete guide in Galapagos islands. Tokyo: Gemstone, pp. 112-114: History of Charles Darwin and Galapagos islands. 2011.

Corbera, J. 2010. La il-lustració a l'obra de Charles Darwin [Illustrating Charles Darwin's work]. L'Atzavara, 19: 27-44.

Cosmos (Catalogue of the Biblioteca Nacional de Espana 2018).

Costa, J. T. 2009. The Darwinian Revelation: Tracing the Origin and Evolution of an Idea. BioScience vol. 59, No. 10 (November): 886-894.

Cracking Comprehension Online. Integra, 2017.

Curtis, V. 2013. Don't go there: The evolution of disgust. Natural History Magazine.

Czigany, Akos. 2015. Darwin Online gallery.

Dahm, Ralf. 2009. Charles Darwin I: Die Anfaenge der Evolutionstheorie. Biospektrum (9 January): 96-97.

Daily Caller.

Darwin in Galapagos. Taiwanese World Journal. 2005, 12.

Darwin Unlimted. Current Biology, vol. 16, issue 22 2006, 11.21

Darwin, C. R. 2009. The expression of the emotions. Joe Cain & Sharon Messenger editors. 2009. [The entire text from the 1890 edition in Darwin Online was used but not acknowledged in the book itself.]

Darwin, C. R. 2009. The origin of species. With an introduction by Julian Huxley. 150th anniversary edition. Signet Classics.

Darwin, C. R. 2018. Charles Darwin: de reis van de Beagle. Amsterdam: L. J. Veen Klassiek.

Darwin, C. R. 2018. Der Ursprung der Arten. translated by Eike Schönfeld with an afterword by Josef H. Reichholf. Stuttgart: Klett-Cotta. (A new translation of the 6th edn)

Darwin, C. R. The Descent of Man. Los Libros de la Catarata (Madrid, 2019)

Darwin's Impact – A Georgia State University Library Exhibit, 2009.

Darwin's Darkest Hour (website/watch online), 2-hour docudrama from NOVA/National Geographic, which aired on PBS on 6 October 2009.

Darwin's Garden. BBC2, Jimmy Doherty. 2009.

David Alexander, The Camel. King Abdulaziz Public Library.

DBH 4 EKI Biologia eta Geologia 4-2 (in Basque), 2016.

DBH 4 EKI Biologia eta Geologia 4-2. [Biology and Geology] (2016). (textbook in Basque).

Delesalle, Nicolas; Portevin, Catherine and Zarachowicz, Weronika eds. 2009. 150 ans après la théorie de l'évolution Charles Darwin dérange encore. Télérama hors série, pp. 1-98.

Derry, J. F. ed. 2010. Darwin in Scotland: Edinburgh, Evolution and Enlightenment. Whittles Publishing.

Detroit Free Press, p. 6. 2009, 02.15

Devon Shiller, The Black Box Grimaces Back: Photographic Evidence in Automated Facial Expression Analysis." Fotografija / Membrana.

Diaries of Darwin's wife to be published online. Telegraph 2007, 03.13

Dorling Kindersley. 2011. Animals Alive: the fight for survival in the wild.

Eckart Voigts-Virchow, Barbara Schaff, Monika Pietrzak-Franger eds. 2013. Reflecting On Darwin: Victorian And Neo-Victorian Responses To Evolutionism. Farnham: Ashgate.

Marchetti, Michael. Ecology, 1ed. W.W. Norton & Company Publishers. 2021.

Farooq, 2015. Undisciplined: Science, Ethnography, and Personhood in the Americas, 1830-1940. New York University Press.

Francis Pryor, Paths to the Past.

Fry, Ben, The Preservation of Favoured Traces (2018).

Fukui Prefectural Dinosaur Museum. 2009. History of the flowering world [in Japanese]. Fukui, pp. 1-97.

Fundación Telefónica, Madrid (exhibition) 2018-2019.

FutureMag (, a 30 mn science programme broadcast on TV channel Arte. 2016, 08.

Gaëtane Maës. 2016. "De Dezallier d'Argenville à Darwin : la question de la couleur dans la représentation des minéraux"

Garnier, Ximena Miranda. 2010. Charles R. Darwin and the development of creativity. Actualidades Investigativas en Educación from the Universidad de Costa Rica 10 No. 3, pp. 1-24.

Geller, Pamela. 2016. The Bioarchaeology of Socio-Sexual Lives - Queering Common Sense about Sex, Gender, and Sexuality. New York: Springer.

Gibson, Sally A. 2009. In Darwin's geological footsteps. Galapagos News, no. 28, pp. 6-7.

Gilbert Lists, du Bois-Reymond's Enigmas, and the Scientific Study of Religion, 2018.

Gingerich, Philip. Rates of Evolution: A Quantitative Synthesis. Cambridge University Press [cover image].

Gliboff, Sander. 2014. Ascent, Descent, and Divergence: Darwin and Haeckel on the Human Family Tree. Konturen 5. http: //

Gomez-Heras, M., McCabe, S.,The weathering of stone-built heritage: a lens through which to read the Anthropocene. Anthropocene.

Google used the microfilm image of Darwin's Notebook B from Darwin Online to create their Darwin Day Google doodle, 2014, 02.12.

Grinkevicius, K., Mikuleviciute, J., Purliene, M. & Firstova, L. 2011. Biologija 7: Vadovelis, pirmoji knyga. Kaunas: Sviesa [Lithuania].

Grisolia, S. 2008. Félix de Azara, Alfred Wallace y Charles Darwin. Biodiversidad (Valencia), pp. 11-24.

Guardian. 2008. Charles Darwin: On the origin of species. The definitive guide to the book that changed the world. The Guardian (Saturday supplement inserted booklet) 34pp.

Hapag Lloyd Cruises. For the new Cruise ships, Hapag is planning a so called "Ocean Academy".

Haydock, Karen. Darwin: 200th anniversary. [Uses Darwin Online images without permission].

Hayman, John. 2009. The Medical Journal of Australia. Med J Aust 2009; 191 (11): 660-663.

Hebrew University of Jerusalem. 2021. MOOC (Massive Open Online Course) on medicine and the Holocaust.

Heiligman, Deborah. 2009. Charles and Emma: The Darwin's leap of faith. New York: Henry Holt and Company, pp. 1-268.

Helvert, Paul van & John van Wyhe. 2021. Darwin: A companion: with iconographies by John van Wyhe. WSP.

Heritage Learning (English Heritage), issue 40, March 2009. With contributions by Skandar Keynes, Sue Johnson, Kim Biddulph, Karen Goldie-Morrison.

Hernández-Laille, Margarita. 2012. Tras los pasos de Charles Darwin por Argentina y Chile. Diario de Lucía Sapiens.

Hessen, Dag O., Lie, Thore & Stenseth, Nils Chr. eds. 2008. Darwin: Verden ble aldri den samme. Oslo: Gyldendal, pp. 1-407.

Historia y cultura 2018. Revista del Museo Nacional de Arqueología, Antropología e Historia del Perú.

Hoch, J. Matthew, Daniel Schneck and Christopher Neufeld. 2016. Ecology and evolution of phenotypic plasticity in the penis and appendages of barnacle, by J. Matthew Hoch, Daniel T. Schneck and Christopher J. Neufeld, Integrative and Comparative Biology.

Holland, Alan. Darwin and the meaning in life. Environmental Values 18 (2009): 503-518.

Holmes, John. 2017. 'Natural History, Evolution and Ecology: The Biological Sciences' in The Routledge Research Companion to Nineteenth-Century British Literature and Science, ed. by John Holmes and Sharon Ruston (Abingdon: Routledge, 2017), 331-56, p. 333.

Holmes, John. The Pre-Raphaelites and Science. Yale.

Hotspot (Informationen des Forum Biodiversität Schweiz), no. 19, Februar 2009. Darwin und die Biodiversität. With contributions by Gregor Klaus, Jurg Stocklin, Reto Nyffeier, Heinz Richner, Walter salzburger, Dieter Ebert, Claus Wedekind, Daniel Prati, Markus Fischer, Bruno Baur, Eva Inderwildi.

Hultgren, Kristin. 2020. Evolution. BIOL 3500.

I, Darwin. National Geographic. 2009.

Inside History (2014, Australian magazine).

Ishikawa, Yuichi. 2013. In the search of Darwin's beetle. Tokyo: Iwasaki Publishing. [in Japanese] 

Johnson, Anna. 'Darwin's Pins: changing attitudes to the authorial trace in a working archive' in Unexpected fame: Conservation approaches to the preparatory object. Proceedings from the International Conference of the Icon Book & Paper Group, Oxford 1–2 October 2018 (London, The Institute of Conservation: 2020). to-the-preparatory-object

Johnston, Elizabeth and Leah Olson. 2015. The Feeling Brain: The Biology and Psychology of Emotions.

Keagan Brewer, 2016. Wonder and Skepticism in the Middle Ages. Routledge.

Kelly, 2012. The 10 most groundbreaking journeys.

Konrad, M. W. 2011. Life on the dock: my introduction to marine biology. Sausalito, California: Science Is Art.

Kort, Pamela, Max Hollein.] 2009. Darwin: Kunst und die Suche nach den Ursprüngen. Schirn Kunsthalle Frankfurt.


Krause, Joachim. 2012. Was Charles Darwin geglaubt hat. Weimar and Eisenach: Wartburg Verlag.

Kyoto University. A Collection of Past University Entrance Exam Papers of Kyoto University (Humanities) (2022 - 2028 editions). Kyogakusha, 56 Iwakura Minamikuwahara-cho Sakyo-ku, Kyoto, 606-0031, Japan.

Laille, 2010. Darwinismo y manuales escolares en Espana e Inglaterra en el siglo XIX (1870-1902).

Lawson, Elizabeth. Primrose (Reaktion books).

Lecointre, Guillaume & Tort, Patrick eds. 2015. Le monde de Darwin. Paris: Editions de La Martinière.

Leesa Rittelmann, "Ins Gesicht geschrieben: Sozialdarwinisten und die photographische Erfassung und Kategorisierung von Menschen' in Streitfall Evolution. Eine Kulturgeschichte.

Leikola, Anto. 2009. Ihminen on osa luontoa, sanoi Darwin. Tiede (Helsinki), 1/2009, pp. 28-31.

Leith Hill Rhododendron Wood. The National Trust. 2009.

Lennox, James, 2016. Instant HPS: Charles Darwin. University of Pittsburgh (online video).

Lidesten, Britt-Marie. 2014. Kolla på djuren! Bi-lagan (Nationellt resurscentrum för biologi och biotechnik, Uppsala) nr. 3: 4-5.

Lindberger, 2012. Entwicklungspsychologie. 7th edn.

Lyons, S. 2011. Evolution: the basics.

MacQuarrie, Life and Death in the Andes: On the Trail of Bandits, Heroes, and Revolutionaries

Macrauchenia Book. Culture Corporation of Calama.

Mallarino et al. 2012. Closely related bird species demonstrate flexibility between beak morphology and underlying developmental programs. PNAS.

Martin Wagner, The Narratology of Observation: Studies in a Technique of European Literary Realism (De Gruyter, 2018).

Marty, Christoph. 2009. Missverständnisse um Darwin. Spektrum der Wissenschaft, Februar 2009, pp. 46-53.

Mason, Daily Rituals (Polish edn)

Mathieu Rivière "Motions of leaves and stems: from growth to potential use" for Physical Biology (IOP Publishing).

max documentary. Secrets of the Universe, produced and directed by Stephen Low. I

McCabe, Thomas. Descent and Logic in Biosystematics.

McInerny, Greg and Stefanie Posavec. 2009. (En)tangled word bank.

Evans, Chris. Celebrating the annus mirabilis. Antiquity 83. 2009, p. 549.

Meyer, Carolyn. 2009. The true adventures of Charley Darwin. Boston and New York: Harcourt, pp. 1-322.

National Museum of Australia. 2008. Charles Darwin: An Australian selection. Canberra: National Museum of Australia Press Ltd., pp. i-xi, 1-131. [With a substantial extract from Kees Rookmaaker's transcription of Darwin's Beagle Diary from Darwin Online (unacknowledged).]

Nature, 9 Oct. 2014 issue, Comment section page 163, print and online.

Neukamm, M., Evolution im Fadenkreuz des Kreationismus. Vandenhoeck & Ruprecht, 2009.

New Zealand Science Review, vol. 66 (3) 2009, the journal of the New Zealand Association of Scientists. "Darwin commemorative issue".

Notes and Records of the Royal Society vol. 61, 2007. The front of the volume reproduces a page from Emma Darwin's diaries, recently published online.

Nucleac Acids Research. 2009.

NUSnews Feb. 2021.

Omnis Cellula, 44-Calaixdesastre. 2010.

On the Face of It: Darwin and the Evolution of Expression. Undark magazine. (3 Nov. 2017)

Opitz, Donald L., Staffan Bergwik, and Brigitte Van Tiggelen eds. 2015. Domesticity in the Making of Modern Science. Basingstoke: Palgrave Macmillan.

Owens, Joana. 2018. HUM385:  Culture, Science, and Technology course packet. LAD Custom Publishing. Atlanta.

Oxendine, Lori J. 2010. "More than kin less than kind", an electronic collage seeking to embrace the organic nature of Darwin's studies and to express the underlying structure of his findings. Genetics in Medicine 12 (2, February): front cover.

Pauli, Margarete. 2009. Vom Ursprung der Arten. Blick (Julius-Maximilians-Universität Würzburg), 1/2009, pp. 19-23.

Pearn, A. M. ed. 2009. A voyage round the world: Charles Darwin and the Beagle collections in the University of Cambridge. Cambridge: CUP.

Pereira, J. N. G. and V. Neves. 2009. Darwin nos Açores. Diário pessoal come comentários. Horta: Observatorio do Mar dos Açores.

Piesse, Jude. 2021. The Ghost in the Garden: In Search of Darwin's Lost Garden.

Pinet, Paul R. 2012. Essential Invitation to Oceanography. Jones & Bartlett Learning.

Pinet, Paul R. 2013. Invitation to oceanography. 6th edn

Plüss, Mathias. 2009. Was Darwin wirklich meinte. Das Magazin, Zürich, 01/2009, pp. 22-31.

Pombo, Olga and Pina, Marco eds. 2012. Em torno de Darwin. Lisbon: Fim de Seculo.

Porr, M. Lives and lines. Integrating molecular genetics, the "origins of modern humans" and Indigenous knowledge. In: A. McGrath & M. A. Jebb eds., Deep histories. Deep time: A transdisciplinary collaboration. Canberra: ANU E Press.

Porter, Duncan M. and Graham, Peter W. 2015. Darwin's sciences; how Charles Darwin voyaged from rocks to worms in his search for facts to explain how the earth, its geological features, and its inhabitants involved. Chichester; Wiley Blackwell. [Darwin Online image mis-attributed and unacknowledged]

Piesse, Jude. 2021. The ghost in the garden: in search of Darwin's lost garden.

Quequicom. (Spanish TV) 2014.

Ranta, Pertti. 2008. Amatööribiologi astui Beagleen. Tiede (Helsinki), 12/2008, pp. 34-41.

Reader, John. 2008. Propitious Esculent: The Potato in World History.

Reader, Simon. Notework: Victorian Literature and Nonlinear Style.

Reidy, Michael S. 2020. Darwin's vertical thinking: Mountains, mobility,and the imagination in 19th-century geology. Centaurus. 62:631-646. [Two colour illustrations used without permission or attribution.]

Reporter, Check Your

Research eu: the magazine of the european research area. Nov. 2008.

Research Horizons (University of Cambridge), issue 9, Summer 2009. Spotlight on Darwin. With contributions by Jim Secord, Alison pearn, Chris Jiggins, David Norman, John Parker, Robert Foley, Marta Mirazon Lahr, Geoff Parks, Jonathan Heeney, Raymond E. Goldstein.

Richards, Evelleen. 2017. Darwin and the Making of Sexual Selection. University of Chicago Press.

Richards, R. A. 2013. The Species Problem: A Conceptual Problem?

Ruban, Alexander V. 2015. Evolution under the sun: optimizing light harvesting in photosynthesis. Journal of Experimental Botany 66 (1): 7-23.

Russell, Hertz, McMillan, Fenton, Maxwell, Haffie, Milsom, Nickle, Ellis. Biology: Exploring the Diversity of Life, 4ce. 2018.

Sachs, Jonathan. The poetics of decline in British Romanticism. Cambridge University Press, c. 2018.

Sacks, Oliver. 2009. Darwin et le secret des fleurs. La Recherche, no. 427: 44-48.

Sánchez J. A & J. López Angarita eds. 2012. Perspectivas en la Ecología de los Arrecifes Coralinos. Universidad de los Andes, Ediciones UniAndes. Bogotá.

Santillan, Oscar. 2012. Text into form.

Schacter, Daniel; Gilbert, Daniel and Wegner, Daniel. 2012. Psychology. First European edn, adapted by Bruce Hood. New York: Palgrave Macmillan. [Images on p. 384]

Loy, James D. and Loy, Kent M. 2010. Emma Darwin: a Victorian life. Gainesville etc.: University Press of Florida. [Images of figures 2, 3, 4, 18, 20].

Fabião, Carlos. 2011. Uma História da Arqueologia Portuguesa [A history of Portuguese Archaeology]. Lisbon: CTT Correios de Portugal. [Images on pp. 102, 103, 104]

Schirn Kunsthalle Frankfurt. 2009. Darwin, Kunst und die Suche nach den Ursprüngen: Eine Einführung in die Ausstellung ab 12 Jahren. Schirn Kunsthalle Frankfurt, 5. Februar bis 14. Juni 2009, pp. 1-36.

Schmitt, C. 2009. Darwin and the Memory of the Human: Evolution, Savages, and South America.

Schneckenburger, Stefan and Omlor, Ralf eds. 2009. Darwins Garten - Evolution entdecken: eine Ausstellung des Verbands Botanischer Gärten zum Darwin-Jahr 2009. Darmstadt and Mainz, Verband Botanischer Gärten.

Scholastic Inc.'s, Classroom Magazine's Division. film 2019.

Schultze, et al. [cover image] 2012. MCP: Molecular & Cellular Proteomics.

Science Friday. 2012. To the bat cave!  

Scruton, Roger. 2014. The Face of God. Continuum International.

Sedgwick Museum of Earth Sciences. (displays) University of Cambridge, 2019.

Sedgwick Museum of Earth Sciences. Darwin in the field. The Sedgwick Museum of Earth Sciences, University of Cambridge. (exhibition since 2009)

Sense of Beauty. Italian Tv Channel 6 part documentary. 2018.

Shuster, Michele; Vigna, Janet; Sinha, Gunjan; Tontonoz, Matthew. 2011. Biology for a changing world. New York: W. H. Freeman and Company. [pp. 290-298: Charles Darwin and Alfred Russel Wallace on the trail of natural selection.]

Siegfried, T. 2009. Darwin's Evolution. ScienceNews

Siegfried, T. 2009. The man who rocked biology to its core. ScienceNewsforStudents.

Siepel, A. 2009. Phylogenomics of primates and their ancestral populations. Genome Research.

Silversea cruise films. 2020.

Skybreak, Ardea. 2006. The science of evolution and the myth of creationism: knowing what's real and why it matters. Chicago: Insight Press, pp. i-viii, 1-338.

Spektrum der Wissenschaft. 02.2009, pp. 46-52.

Spektrum der Wissenschsft online video about wolvess/dogs, 2018.

Stanford continuing studies Spring 2017 courses brochure (104pp).

Stangroom, Einstein's Twin. Elwin Street Productions | Modern Books.

Steinmüller, Angela and Karlheinz Steinmüller. 2008. Darwins Welt: Aus dem Leben eines unfreiwilligen Revolutionärs.

Stöcklin, J. 2009. Darwin and the Plants of the Galápagos-Islands. Bauhinia 21, pp. 33-48.

Stone, Irving. 2011. The Origin (in Polish).

Straits Times, Nov. 2020.

Stringham et al. 2012. Divergence, Convergence, and the Ancestry of Feral Populations in the Domestic Rock Pigeon, Current Biology, doi:10.1016/j.cub.2011.12.045

Tanghe, Koen B. 2019. On The Origin of Species: The story of Darwin's title. Notes & records of the Royal Society 73, 83-100.

Takako, Hirakawa. 2010. [Wonder of the Galapagos] (In Japanese) Japan Association for Galapagos.

Taylor, Bayard. 2008. The late great ape debate. Cincinnati, Ohio: Standard Publishing, pp. 1-208.

Taylor, James. 2008. The voyage of the Beagle: Darwin's extraordinary adventure aboard FitzRoy's famous survey ship. London: Conway, pp. 1-192. (unacknowledged maps reproduced pp. 16-17, 129).

Ten Brinke & S. Porter. 2012. Face to Face: An Integrated Theory of the Evolutionary Struggle Between Voluntary and Involuntary Influences on Human Facial Communication. Perspectives on Psychological Science

Tethys Himalayan Fossil Museum of Evolution in Kasauli, Himachal Pradesh, India.

Text and images etched into a sheet of patinated brass. bench back rests at Crystal Palace Park (London Borough of Bromley)

The Times Literary Supplement April 25, 2008; p. 3.

Thompson, Ken. 2018. Darwin's Most Wonderful Plants. Profile Books.

Trigo, Nekae ed. Intercostaliteraria (a magazine about literature and philosophy that is published in Spain)

Trontelj, Peter. 2009. Charles Darwin, evolucijska teorija in ptice. Svet ptic, no.2, pp. 36-37.

Turney, Jon. 2009. Darwin now. Manchester: British Council, pp. 1-60.

TVBrasil. 2016, TV show called "Expedições" (expeditions), displayed on TVBrasil (a public tv station from Brasil).

Twenty First Century Science. Oxford University Press, 2011.

Vala, Filipa ed. 2009. Darwin en Cabo Verde. Lisboa: Gulbenkian, pp. 1-88.

Vav Vranken, David & Weiss, Gregory. 2013. Introduction to bioorganic chemistry and chemical biology. New York and London: Garland Science.

VDI nachrichten (a German weekly for engineers). 2019, 12.

Villavicencio, C. 2017. Dimensiones Paralelas.

Villavicencio, Cristian. Bilbao exhibition catalogue, 2017.

WakeWood Productions Ltd. Online film for Leith Hill Place,  2018.

Wallace, A. R. 2018. Het Maleisische eilandenrijk van Alfred Russel Wallace. Amsterdam: L. J. Veen Klassiek.

Warne, Kennedy. 2009. Irritable in Aotearoa: Darwin & the Barbarians. New Zealand Geographic, no. 95, January-February 2009, pp. 58-69.

Wartenberg, Henry von. 2010. Charles Darwin at southern south: on his tracks two centuries later. Tigre: Tripleve Editores.

Wildlife; Galapagos Iguana. 12 November, 2012 TV: NHK (Japan Broadcasting Corporation)

Williams, G. 2013. Naturalists at Sea: Scientific Travellers from Dampier to Darwin. Yale.

Winston, Robert. 2009. Evolution Revolution.

Wisconsin Magazine of History, Nov. 2020.

Wood, Samuel E.; Green Wood, Ellen; Boyd, Denise and Hétu, Francine eds. 2009. L'Univers de la Psychologie. Québec: Editions du Renouveau Pedagogique, pp. 1-444.

Wyhe, John van. Darwin. (Andre Deutsch, 2008; National Geographic, Oct. 2009).

Wyhe, John van ed. 2009. Charles Darwin's Shorter Publications 1829-1883. Cambridge University Press.

Wyhe, John van. 2009. Darwin in Cambridge. (Christ's College, 2009).

Wyhe, John van. 2009. Darwin vs God? BBC History Magazine, 10 (1), January 2009, pp. 26-31.

Wyhe, John van. 2012. Correcting the Darwin myths. In The Origin of Darwinism: Charles Darwin and The Origin of Species. Guardian Books.

Wyhe, John van ed. 2012-. Wallace Online (

Wyhe, John van. 2013. Dispelling the Darkness: Voyage in the Malay Archipelago and the discovery of evolution by Wallace and Darwin. WSP.

Wyhe, John van. 2014. Charles Darwin in Cambridge: The most joyful years. WSP.

Wyhe, John van. 2018. Darwin: The man, his great voyage, and his theory of evolution. NHM, A. Deutsch. (translated into: French, Spanish, Czech, Chinese, Thai, Vietnamese and Italian)

Wyhe, John van. 2020. Charles Darwin: The compact guide. Welbeck.

Wyhe, John van ed. 2020. On the origin of species. The science classic. With an introduction by John van Wyhe. Capstone Classics. [Introduction available in Darwin Online: Text]

Wyhe, John van & Christine Chua. 2021. Charles Darwin: Justice of the peace. The complete records (1857-1882). PDF

Wyhe, John van. 2021. Charles Darwin on Tahiti. Bulletin de la Société des Études Océaniennes, no. 355 (Sept./Dec.): 106-170. [English & French] PDF

Wyhe, John van. 2022. Charles Darwin: the man, his great voyage, and his theory of evolution. Pioneers of Science. New York: Rosen.

Wynn-Jones, Elizabeth and Byers, Jenny. 2009. The most curious tourist. British and Commonwealth Society of Rio de Janeiro, Yearbook, 2009, pp. 4-6, 19-23, 49, 88.

Zarimis, Maria. 2015. Darwin's footprint: cultural perspectives on evolution in Greece (1880-1930s).

Zordan, Dr. Luca, Zwischen Mythos und Wissenschaft. Ökologisierung in der Zusammenarbeit von Künstlern und Wissenschaftlern im 19. Jahrhundert: Kretschmers und Brehms Illustrirtes Thierleben, transcript Verlag.

Putten, Kees van, Trees, Coral, and Seaweed: An Interpretation of Sketches Found in Darwin's Papers. Journal of the History of Biology vol. 53, 5-44. 2020.

O pensamento evolutivo antes de Darwin" ("The evolutionary thinking before Darwin"), in the Brazilian journal Genética na Escola. 2021.

Conor Barry, Darwin's Theory of the Origin of Species by Natural Selection and the Research Community of The Voyage of the Beagle. International Society for the History, Philosophy and Social Studies of Biology.

Daniel Schacter, Daniel Gilbert, Mathew Nock, Daniel Wegner, Psychology 6e, Introducing Psychology 6e and Psychology Canadian 6e Product Family, 6e. Bedford, Freeman & Worth Publishing Group, LLC d/b/a Macmillan Learning.

James G. Lennox, Accentuate the negative: Locating possibility in Darwin's 'long argument'. Studies in History and Philosophy of Science 87 (2021) 147-157.

Planet Explorers: The Advanced Rover Exploring Mars. BBC Earth. 2021.

Kay Harel, Darwin's Love of Life: A Singular Case of Biophilia. Columbia University Press, 2022.

Darwin - on the origin of species. 'Magic moments of mankind' series. Müller und Schindler. 2022. Cover art.

Project with ExpertEd on the Voyage of the Beagle, 2022.

Scott et al. Biological Sciences: Exploring the Science of Life, OUP, 2022.

James J. A. Blair, 2022. Salvaging empire: Sovereignty, natural resources and environmental science in the South Atlantic. Cornell University Press.

Kay Harel, Darwin's dogs. ReVista, the Harvard Review of Latin America, 2023.

Justine Blau. K. Verlag, 2023.

W. S. Armbruster, Plant Behavioral Ecology: New insights into the adaptive significance of rapid floral movements. Dispatches. Current Biology, 2023.

Tom McAvoy, Science supports faith: A book for the non-scientist.

David Gordon, Naturalism, and Theism: The Implications of Evolution for Metaphysics. Lexington Books, 2023.

Joy Hakim, Discovering Life's Story, Volume 2: The Evolution of an Idea. 2023.

John Dunn, [Article on Darwin in Chiloe, quoting from Rookmaaker's transcription of the Beagle diary]. Firmament. 2023.

Nancy Campbell, Nature Tales for Winter Nights. Elliott & Thompson.

Exhibition catalogue: 'Simple, Complex, Alive. How matter became life' Deutsches Museum, München. 2023.

Learning Evolutionary Biology through Quizzes, Kadokawa Co., Ltd. (Japan) 2023.

Matteo Mameli, Why Human Nature Matters. Bloomsbury Academic.

Sydney Srinivas, Charles Darwin. Prism, India, 2022.

Rasheed Tazudeen, Modernism's Inhuman Worlds. Cornell University Press, 2024.

Gary Prost, Geologic Tours of the World – South America’s Natural Wonders. Taylor and Francis/CRC Press, 2024.

Michael Taylor, Impossible Monsters: Dinosaurs, Darwin and the War between Science and Religion (Bodley Head, 2024).

Lindell Bromham, Combining molecular, macroevolutionary and macroecological perspectives on the generation of diversity. In Catherine Piechel ed., Cold Spring Harbor Perspective on Speciation.

Exhibition catalogue on the Austrian actor Hugo Thimig, 2024.

Audrey Rieber, L'histoire depuis l'art. ENS Editions (Lyon).

Christopher Jones (12.2203) has made the most detailed map ever of Darwin's voyage of the Beagle using FitzRoy's Narrative and Kees Rookmaaker's day-by-day itinerary of the voyage and his new transcription of the Beagle Diary- all in Darwin Online.

Kate Holterhoff, Speculation and the Darwinian Method in British Romance Fictions, 1859-1914.  

Joyce Dixon, PhD thesis on the publication of Werner's Nomenclature of Colours (1821). University of Edinburgh.

Peter Greenaway film, "The Tower Stories".

Jelle Reumer, a history of paleontology in the 18th and 19th centuries.

Graphic novel on Darwin's life, written and illustrated by Camille Van Belle and Adrien Miqueu. Alisio, a French imprint of Leduc-Albin Michel Group.

Jon Rouco trans. Charles Darwin. Autobiografía Completa: Con todas las omisiones restauradas. (Spanish translation of Autobiography with omissions restored.) 2024.

Darwin, On the Origin of Species, 6th edition. Portuguese translation. Brazil: Companhia das Letras. 2024.

Edmund Higgins, The Neuroscience of Clinical Psychiatry, 4th ed. Wolters Kluwer, 2024.

Hermes Foundation, Ginza Maison Hermes Le Forum, an art gallery, Tokyo.

Martinez-Reina, Charles Darwin, a philatelic-didactice biography. Resonance. September 2022.

van der Gulik, P. T. S., Hoff, W. D., & Speijer, D. (2024). The contours of evolution: In defence of Darwin's Tree of Life paradigm. BioEssays, 46.


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