Origin of Species Variorum

Comparison with 1869

much older fishes, of which the affinities are as yet imperfectly known, are really teleostean. Assuming, however, that the whole of them did appear, as Agassiz maintains, at the commencement of the chalk formation, the fact would certainly be highly remarkable; but I cannot see that it would be an insuperable objection to these views, unless it could likewise be shown that the species of this group appeared suddenly and simultaneously throughout the world at this same period. It is almost superfluous to remark that hardly any fossil-fish are known from south of the equator; and by running through Pictets Palæontology it will be seen that very few species are known from several formations in Europe. Some few families of fish now have a confined range; the teleostean fish might formerly have had a similarly confined range, and after having been largely developed in some one sea, might have spread widely. Nor have we any right to suppose that the seas of the world have always been so freely open from south to north as they are at present. Even at this day, if the Malay Archipelago were converted into land, the tropical parts of the Indian Ocean would form a large and perfectly enclosed basin, in which any great group of marine animals might be multiplied; and here they would remain confined, until some of the species became adapted to a cooler climate, and were enabled to double the southern capes of Africa or Australia, and thus reach other and distant seas.

From these considerations, from our ignorance of the geology of other countries beyond the confines of Europe and the United States, and from the revolution in our palæontological knowledge effected by the discoveries of .. the last dozen years, .. it seems to me to be about as rash .. to dogmatize on the succession of organic

much older fishes, of which the affinities are as yet imperfectly known, are really teleostean. Assuming, however, that the whole of them did appear, as Agassiz believes, at the commencement of the chalk formation, the fact would certainly be highly remarkable; but I cannot see that it would be an insuperable difficulty on my theory, unless it could likewise be shown that the species of this group appeared suddenly and simultaneously throughout the world at this same period. It is almost superfluous to remark that hardly any fossil-fish are known from south of the equator; and by running through Pictet's Palæontology it will be seen that very few species are known from several formations in Europe. Some few families of fish now have a confined range; the teleostean fish might formerly have had a similarly confined range, and after having been largely developed in some one sea, might have spread widely. Nor have we any right to suppose that the seas of the world have always been so freely open from south to north as they are at present. Even at this day, if the Malay Archipelago were converted into land, the tropical parts of the Indian Ocean would form a large and perfectly enclosed basin, in which any great group of marine animals might be multiplied; and here they would remain confined, until some of the species became adapted to a cooler climate, and were enabled to double the southern capes of Africa or Australia, and thus reach other and distant seas.

From these and similar considerations, but chiefly from our ignorance of the geology of other countries beyond the confines of Europe and the United States; and from the revolution in our palæontological ideas on many points, which the discoveries of even the last dozen years have effected, it seems to me to be about as rash in us to dogmatize on the succession of organic