Origin of Species Variorum

Comparison with 1866

indispensable for the preservation of all the transitional gradations between any two or more species. If such gradations were not all fully preserved, transitional varieties would merely appear as so many distinct species. It is also probable that each great period of subsidence would be interrupted by oscillations of level, and that slight climatal changes would intervene during such lengthy periods; and in these cases the inhabitants of the archipelago would .. migrate, and no closely consecutive record of their modifications could be preserved in any one formation.

Very many of the marine inhabitants of the archipelago now range thousands of miles beyond its confines; and analogy plainly leads to the belief that it would be chiefly these far-ranging species, though only some of them, which would oftenest produce new varieties; and the varieties would at first generally be local or confined to one place, but if possessed of any decided advantage, or when further modified and improved, they would slowly spread and supplant their parent-forms. When such varieties returned to their ancient homes, as they would differ from their former state, in a nearly uniform, though perhaps extremely slight degree, and as they would be found embedded in slightly different sub-stages of the same formation, they would, according to the principles followed by many palæontologists, be ranked as new and distinct species.

If then there be some degree of truth in these remarks, we have no right to expect to find, in our geological formations, an infinite number of those fine transitional forms which, on my theory, .. have connected all the past and present species of the same group into one long and branching chain of life. We ought only to look for a few links, and such assuredly we do find— some more distantly, some more closely, related to each other; and these links, let them be ever so close, if found in different stages of the same formation, would, by many palæontologists,

indispensable for the preservation of all the transitional gradations between any two or more species. If such gradations were not fully preserved, transitional varieties would merely appear as so many distinct species. It is, also, probable that each great period of subsidence would be interrupted by oscillations of level, and that slight climatal changes would intervene during such lengthy periods; and in these cases the inhabitants of the archipelago would have to migrate, and no closely consecutive record of their modifications could be preserved in any one formation.

Very many of the marine inhabitants of the archipelago now range thousands of miles beyond its confines; and analogy leads me to believe that it would be chiefly these far-ranging species which would oftenest produce new varieties; and the varieties would at first generally be local or confined to one place, but if possessed of any decided advantage, or when further modified and improved, they would slowly spread and supplant their parent-forms. When such varieties returned to their ancient homes, as they would differ from their former state, in a nearly uniform, though perhaps extremely slight degree, they would, according to the principles followed by many palæontologists, be ranked as new and distinct species.

If then, there be some degree of truth in these remarks, we have no right to expect to find in our geological formations, an infinite number of those fine transitional forms, which on my theory assuredly have connected all the past and present species of the same group into one long and branching chain of life. We ought only to look for a few links, some more closely, some more distantly related to each other; and these links, let them be ever so close, if found in different stages of the same formation, would, by most palæonto- logists,