Origin of Species Variorum

Comparison with 1866

.. elapsed between our consecutive formations,— longer perhaps in many cases than the time required for the accumulation of each formation. These intervals will have given time for the multiplication of species from some one or some few parent-forms; and in the succeeding formation such groups of species will appear as if suddenly created.

I may here recall a remark formerly made, namely, that it might require a long succession of ages to adapt an organism to some new and peculiar line of life, for instance, to fly through the air; and consequently that the transitional forms would often long remain confined to some one region; but that, when this adaptation had once been effected, and a few species had thus acquired a great advantage over other organisms, a comparatively short time would be necessary to produce many divergent forms, which would be able to spread rapidly and widely throughout the world. Professor Pictet, in his excellent Review of this work, in commenting on early transitional forms, and taking birds as an illustration, cannot see how the successive modifications of the anterior limbs of a supposed prototype could possibly have been of any advantage. But look at the penguins of the Southern Ocean; have not these birds their front limbs in this precise intermediate state of "neither true arms nor true wings"? Yet these birds hold their place victoriously in the battle for life; for they exist in infinite numbers and of many kinds. I do not sup- pose that we here see the real transitional grade through which the wings of birds have passed; but what special difficulty is there in believing that it might profit the modified descendants of the penguin, first to become enabled to flap along the surface of the sea like the logger-headed duck, and ultimately to rise from its surface and glide through the air?

I will now give a few examples to illustrate the foregoing remarks, and to show how liable we are to error in supposing that whole groups of species have suddenly been produced. Even in so short an interval as that between the first and second editions of Pictet's great work on Palæontology, published in 1844-46 and in 1853-57, the conclusions on the first appearance and disappearance of several groups of animals have been considerably modified; and a third edition would require still further changes. I may recall the well-known fact that in geological treatises, published not many years ago, the whole class of mammals was always spoken of as having abruptly come in at the commencement of the tertiary series. And now one of the richest known accumulations of fossil mammals, for its thickness, belongs to the middle of the secondary series; and .. true mammals have been discovered in the new red sandstone at nearly the commencement of this great series. Cuvier used to urge that no monkey occurred in any tertiary stratum; but now extinct species have been discovered in India, South America, and in Europe, as far back as the miocene stage.

Text in this page (from paragraph 4000, sentence 400 to paragraph 4000, sentence 500, word 38) is not present in 1866

probably elapsed between our consecutive formations,— longer perhaps in some cases than the time required for the accumulation of each formation. These intervals will have given time for the multiplication of species from some one or some few parent-forms; and in the succeeding formation such species will appear as if suddenly created.

I may here recall a remark formerly made, namely that it might require a long succession of ages to adapt an organism to some new and peculiar line of life, for instance to fly through the air; but that when this had been effected, and a few species had thus acquired a great advantage over other organisms, a comparatively short time would be necessary to produce many divergent forms, which would be able to spread rapidly and widely throughout the world. ↑

I will now give a few examples to illustrate these remarks; and to show how liable we are to error in supposing that whole groups of species have suddenly been produced. ↑ I may recall the well-known fact that in geological treatises, published not many years ago, the great class of mammals was always spoken of as having abruptly come in at the commencement of the tertiary series. And now one of the richest known accumulations of fossil mammals belongs to the middle of the secondary series; and one true mammal has been discovered in the new red sandstone at nearly the commencement of this great series. Cuvier used to urge that no monkey occurred in any tertiary stratum; but now extinct species have been discovered in India, South America, and in Europe even as far back as the eocene stage. The most striking case, however, is that of the Whale family; as these animals have huge bones, are marine, and range over the world, the fact of not a single bone of a whale having been discovered in