| Comparison with 1869 | 
  | 
formations, by the physical conditions of the ancient areas having remained nearly the same.  Let it be remembered that the forms of life, at least those inhabiting the sea, have changed almost simultaneously throughout the world, and therefore under the most different climates and conditions.  Consider the prodigious 
vicissitudes of climate during the pleistocene period, which includes the whole glacial 
 epoch, | epoch, 1869 1872 |  | period, 1859 1860 1861 1866 |  
  
and note how little the specific forms of the inhabitants of the sea have been affected.  | 
 | 
 On the theory of descent, the full meaning of the 
 ..| ..... 1869 1872 |  | fact of 1859 1860 1861 1866 |  
  
fossil remains from closely consecutive 
 formations | formations 1869 1872 |  | formations, 1859 1860 1861 1866 |  
  
 being closely related, though ranked as distinct species, | being closely related, though ranked as distinct species, 1869 1872 |  
| though ranked as distinct species, being closely related, 1859 1860 1861 1866 |  
  
is obvious.  As the accumulation of each formation has often been interrupted, and as long blank intervals have intervened between successive formations, we ought not to expect to find, as I attempted to show in the last chapter, in any one or 
 in any two | in any two 1866 1869 1872 |  
| two 1859 1860 1861 |  
  
formations 
all the intermediate varieties between the species which appeared at the commencement and close of these 
 periods: | periods: 1869 1872 |  | periods; 1859 1860 1861 1866 |  
  
but we ought to find after intervals, very long as measured by years, but only moderately long as measured geologically, closely allied forms, or, as they have been called by some authors, representative species; and these 
 assuredly we | assuredly we 1866 1869 1872 |  | we assuredly 1859 1860 1861 |  
  
do find.  We find, in short, such evidence of the slow and scarcely sensible mutation 
of specific forms, as we have 
 the | the 1869 1872 |  | a just 1859 1860 1861 1866 |  
  
right to 
 expect. | expect. 1866 1869 1872 |  | expect 1859 1860 1861 |  
  
 ..| ..... 1866 1869 1872 |  | to 1859 1860 1861 |  
  
 ..| ..... 1866 1869 1872 |  | find. 1859 1860 1861 |  
  
 | 
 
On  
the 
 
State
 | 
State
 1861 1869 |  | 
state
 1859 1860 |  | 
State
 1866 1872 |  
   of  
Development  
of  
Ancient 
 
compared with Living Forms.
 | 
compared with Living Forms.
 1869 |  
| 
Forms
.—
 1859 1860 |  
| 
compared with Living Forms.
—
 1861 |  
| 
compared with Living Forms.  1866 |  
| 
compared with Living Forms
.  1872 |  
  | 
| Text in this page (from  paragraph 3700, sentence 010 to  paragraph 3700, sentence 300, word 6) is not present in 1869 | 
 
  
  
formations, by the physical conditions of the ancient areas having remained nearly the same.  Let it be remembered that the forms of life, at least those inhabiting the sea, have changed almost simultaneously throughout the world, and therefore under the most different climates and conditions.  Consider the 
 prodigious | prodigious 1859 1861 1866 1869 1872 |  | pro- digious 1860 |  
  
vicissitudes of climate during the pleistocene period, which includes the whole glacial 
 period, | period, 1859 1860 1861 1866 |  | epoch, 1869 1872 |  
  
and note how little the specific forms of the inhabitants of the sea have been affected.  | 
 | 
 On the theory of descent, the full meaning of the 
 fact of | fact of 1859 1860 1861 1866 |  fact of 1869 1872 |  
  
fossil remains from closely consecutive 
 formations, | formations, 1859 1860 1861 1866 |  | formations 1869 1872 |  
  
 though ranked as distinct species, being closely related, | though ranked as distinct species, being closely related, 1859 1860 1861 1866 |  
| being closely related, though ranked as distinct species, 1869 1872 |  
  
is obvious.  As the accumulation of each formation has often been interrupted, and as long blank intervals have intervened between successive formations, we ought not to expect to find, as I attempted to show in the last chapter, in any one or 
 two | two 1859 1860 1861 |  
| in any two 1866 1869 1872 |  
  
 formations | formations 1859 1860 1861 1866 1869 |  | formations, 1872 |  
  
all the intermediate varieties between the species which appeared at the commencement and close of these 
 periods; | periods; 1859 1860 1861 1866 |  | periods: 1869 1872 |  
  
but we ought to find after intervals, very long as measured by years, but only moderately long as measured geologically, closely allied forms, or, as they have been called by some authors, representative species; and these 
 we assuredly | we assuredly 1859 1860 1861 |  | assuredly we 1866 1869 1872 |  
  
do find.  We find, in short, such evidence of the slow and scarcely sensible 
 mutation | mutation 1859 1860 1861 1866 1869 |  | mutations 1872 |  
  
of specific forms, as we have 
 a just | a just 1859 1860 1861 1866 |  | the 1869 1872 |  
  
right to 
 expect | expect 1859 1860 1861 |  | expect. 1866 1869 1872 |  
  
 to | to 1859 1860 1861 |  to 1866 1869 1872 |  
  
 find. | find. 1859 1860 1861 |  find. 1866 1869 1872 |  
  
 | 
 
On
 | 
On
 1859 1860 1861 1869 |  | 
On
 1866 1872 |  
  
 
the
 | 
the
 1859 1860 1861 1869 |  | 
the
 1866 1872 |  
  
 
state
 | 
state
 1859 1860 |  | 
State
 1861 1869 |  | 
State
 1866 1872 |  
  
 
of
 | 
of
 1859 1860 1861 1869 |  | 
of
 1866 1872 |  
  
 
Development
 | 
Development
 1859 1860 1861 1869 |  | 
Development
 1866 1872 |  
  
 
of
 | 
of
 1859 1860 1861 1869 |  | 
of
 1866 1872 |  
  
 
Ancient
 | 
Ancient
 1859 1860 1861 1869 |  | 
Ancient
 1866 1872 |  
  
 
Forms
.—
 | 
Forms
.—
 1859 1860 |  
| 
compared with Living Forms.
—
 1861 |  
| 
compared with Living Forms.  1866 |  
| 
compared with Living Forms.
 1869 |  
| 
compared with Living Forms
.  1872 |  
  
↑| 2 blocks not present in  1859 1860; present in  1861 1866 1869 1872 |  |  We have seen in the fourth chapter that the degree of differentiation and specialisation of the parts in 
all 
organic beings, when arrived at 
maturity, is the best standard, as yet suggested, of their degree of perfection or highness. 
We have also seen that, 
as the specialisation of parts and organs 
is an advantage to each being, so natural selection will tend 
to render the organisation of each being more specialised and perfect, and in this sense higher; not but that it may and will 
leave many creatures with simple and unimproved structures fitted for simple conditions of life, and in some cases will even degrade or simplify the organisation, yet leaving such degraded beings better fitted for their new walks of life. 
 |  
   There has been much discussion whether recent forms are more highly developed than ancient.  I will not here enter on this subject, for naturalists have not as yet defined to each other's satisfaction what is meant by high and low forms. ↑| 1 blocks not present in  1859 1860 1861 1866 1869 1872; present in  |  | 
 |  
   
But in one particular sense the 
 |