Origin of Species Variorum

Comparison with 1872

(though I do not believe in this doctrine) by formerly continuous land, .. and the Cape de Verde Islands from Africa; .. such colonists would be liable to modification,— the principle of inheritance still betraying their original birthplace.

Many analogous facts could be given: indeed it is an almost universal rule that the endemic productions of islands are related to those of the nearest continent, or of the nearest large island. The exceptions are few, and most of them can be explained. Thus although Kerguelen Land stands nearer to Africa than to America, the plants are related, and that very closely, as we know from Dr. Hooker's account, to those of America: but on the view that this island has been mainly stocked by seeds brought with earth and stones on icebergs, drifted by the prevailing currents, this anomaly disappears. New Zealand in its endemic plants is much more closely related to Australia, the nearest mainland, than to any other region: and this is what might have been expected; but it is also plainly related to South America, which, although the next nearest continent, is so enormously remote, that the fact becomes an anomaly. But this difficulty partially disappears on the view that .. New Zealand, South America, and the other southern lands have been stocked in part from a nearly intermediate though distant point, namely from the antarctic islands, when they were clothed with vegetation, during a warmer tertiary period, before the commencement of the last Glacial period. The affinity, which though feeble, I am assured by Dr. Hooker is real, between the flora of the south-western corner of Australia and of the Cape of Good Hope, is a far more remarkable case; ... but this affinity is confined to the plants, and will, no doubt, some day be explained.

The same law which has determined the relationship between the inhabitants of islands and

by formerly continuous land, from America; and the Cape de Verde Islands from Africa; and that such colonists would be liable to modification;— the principle of inheritance still betraying their original birthplace.

Many analogous facts could be given: indeed it is an almost universal rule that the endemic productions of islands are related to those of the nearest continent, or of other near islands. The exceptions are few, and most of them can be explained. Thus the plants of Kerguelen Land, though standing nearer to Africa than to America, are related, and that very closely, as we know from Dr. Hooker's account, to those of America: but on the view that this island has been mainly stocked by seeds brought with earth and stones on icebergs, drifted by the prevailing currents, this anomaly disappears. New Zealand in its endemic plants is much more closely related to Australia, the nearest mainland, than to any other region: and this is what might have been expected; but it is also plainly related to South America, which, although the next nearest continent, is so enormously remote, that the fact becomes an anomaly. But this difficulty almost disappears on the view that both New Zealand, South America, and other southern lands were long ago partially stocked from a nearly intermediate though distant point, namely from the antarctic islands, when they were clothed with vegetation, before the commencement of the Glacial period. The affinity, which, though feeble, I am assured by Dr. Hooker is real, between the flora of the south-western corner of Australia and of the Cape of Good Hope, is a far more remarkable case, and is at present inexplicable: but this affinity is confined to the plants, and will, I do not doubt, be some day explained.

The law which causes the inhabitants of an archi- pelago, though specifically distinct, to be closely allied to those of