Origin of Species Variorum

Comparison with 1866

orders, under still fewer classes, and all in one great natural system. As showing how few the higher groups are in number, and how widely spread they are throughout the world, the fact is striking, that the discovery of Australia has not added an insect belonging to a new class; and that in the vegetable kingdom, as I learn from Dr. Hooker, it has added only two or three families of small size.

In the chapter on Geological Succession I attempted to show, on the principle of each group having generally diverged much in character during the long-continued process of modification, how it is that the more ancient forms of life often present characters in some slight degree intermediate between existing groups. A few old and intermediate parent-forms having occasionally transmitted to the present day descendants but little modified, will give to us our so-called osculant or aberrant groups. The more aberrant any form is, the greater must be the number of connecting forms which on my theory have been exterminated and utterly lost. And we have some evidence of aberrant forms having suffered severely from extinction, for they are generally represented by extremely few species; and such species as do occur are generally very distinct from each other, which again implies extinction. The genera Ornithorhynchus and Lepidosiren, for example, would not have been less aberrant had each been represented by a dozen species instead of by a single one; but such richness in species, as I find after some investigation, does not commonly fall to the lot of aberrant genera. We can, I think, account for this fact only by looking at aberrant forms as failing groups conquered by more successful competitors, with a few members preserved by some unusual coincidence of favourable circumstances.

Mr. Waterhouse has remarked that, when a member

orders, under still fewer classes, and all in one great natural system. As showing how few the higher groups are in number, and how widely spread they are throughout the world, the fact is striking, that the discovery of Australia has not added a single insect belonging to a new order; and that in the vegetable kingdom, as I learn from Dr. Hooker, it has added only two or three orders of small size.

In the chapter on geological succession I attempted to show, on the principle of each group having generally diverged much in character during the long-continued process of modification, how it is that the more ancient forms of life often present characters in some slight degree intermediate between existing groups. A few old and intermediate parent-forms having occasionally transmitted to the present day descendants but little modified, will give to us our so-called osculant or aberrant groups. The more aberrant any form is, the greater must be the number of connecting forms which on my theory have been exterminated and utterly lost. And we have some evidence of aberrant forms having suffered severely from extinction, for they are generally represented by extremely few species; and such species as do occur are generally very distinct from each other, which again implies extinction. The genera Ornithorhynchus and Lepidosiren, for example, would not have been less aberrant had each been represented by a dozen species instead of by a single one; but such richness in species, as I find after some investigation, does not commonly fall to the lot of aberrant genera. We can, I think, account for this fact only by looking at aberrant forms as failing groups conquered by more successful competitors, with a few members preserved by some unusual coincidence of favourable circumstances.

Mr. Waterhouse has remarked that, when a member