Origin of Species Variorum

Comparison with 1861

together by .. long, but broken chains of affinities. Extinction has only separated groups: it has by no means made them; for if every form which has ever lived on this earth were suddenly to reappear, though it would be quite impossible to give definitions by which each group could be distinguished from other groups, as all would blend together by steps as fine as those between existing varieties, nevertheless a natural classification, or at least a natural arrangement, would be possible. We shall see this by turning to the diagram: the letters, A to L, may represent eleven Silurian genera, some of which have produced large groups of modified descendants. Every intermediate link between these eleven genera and their primordial parent, and every intermediate link in each branch and sub-branch of their descendants, may be supposed to be still alive; and the links to be as fine as those between the finest varieties. In this case it would be quite impossible to give any definition by which the several members of the several groups could be distinguished from their more immediate parents; or these parents from their ancient and unknown progenitor. Yet the natural arrangement in the diagram would still hold good; and, on the principle of inheritance, all the forms descended from A, or from I, would have something in common. In a tree we can specify this or that branch, though at the actual fork the two unite and blend together. We could not, as I have said, define the several groups; but we could pick out types, or forms, representing most of the characters of each group, whether large or small, and thus give a general idea of the value of the differences between them. This is what we should be driven to, if we were ever to succeed in collecting all the forms in any class which have lived throughout all time and space. We shall assuredly never succeed in making

together by a long, but broken, chain of affinities. Extinction has only separated groups: it has by no means made them; for if every form which has ever lived on this earth were suddenly to reappear, though it would be quite impossible to give definitions by which each group could be distinguished from other groups, as all would blend together by steps as fine as those between the finest existing varieties, nevertheless a natural classification, or at least a natural arrangement, would be possible. We shall see this by turning to the diagram: the letters, A to L, may represent eleven Silurian genera, some of which have produced large groups of modified descendants. Every intermediate link between these eleven genera and their primordial parent, and every intermediate link in each branch and sub-branch of their descendants, may be supposed to be still alive; and the links to be as fine as those between the finest varieties. In this case it would be quite impossible to give any definition by which the several members of the several groups could be distinguished from their more immediate parents; or these parents from their ancient and unknown progenitor. Yet the natural arrangement in the diagram would still hold good; and, on the principle of inheritance, all the forms descended from A, or from I, would have something in common. In a tree we can specify this or that branch, though at the actual fork the two unite and blend together. We could not, as I have said, define the several groups; but we could pick out types, or forms, representing most of the characters of each group, whether large or small, and thus give a general idea of the value of the differences between them. This is what we should be driven to, if we were ever to succeed in collecting all the forms in any class which have lived throughout all time and space. We shall certainly never succeed in making