Origin of Species Variorum

Comparison with 1869

rule, that a rudimentary part .. is of greater size relatively to the adjoining parts in the embryo, than in the adult; so that the organ at this early age is less rudimentary, or even cannot be said to be in any degree rudimentary. Hence .. rudimentary organs in the adult are often said to have retained their embryonic condition.

I have now given the leading facts with respect to rudimentary organs. In reflecting on them, every one must be struck with astonishment: for the same reasoning power which tells us .. that most parts and organs are exquisitely adapted for certain purposes, tells us with equal plainness that these rudimentary or atrophied organs are imperfect and useless. In works on natural history, rudimentary organs are generally said to have been created "for the sake of symmetry," or in order "to complete the scheme of nature." .. .. .. .. .. .. .. .. .. .. .. .. .. But this is not an explanation, merely a re-statement of the fact. Nor is it consistent with itself: thus the boa-constrictor has rudiments of hind-limbs and of a pelvis, and if it be said that these bones have been retained "to complete the scheme of nature," why, as Professor Weismann asks, have they not been retained by other snakes, which do not possess even a vestige of these same bones? What would be thought of an astronomer, who maintained that the satellites revolve in elliptic courses round their planets "for the sake of symmetry," because the planets thus revolve round the sun? An eminent physiologist accounts for the presence of rudimentary organs, by supposing that they serve to excrete matter in excess, or matter injurious to the system; but can we suppose that the minute papilla, which often represent the pistil in male flowers, and which is formed .. of mere cellular tissue, can thus act? Can we suppose that ... rudimentary teeth, which are subsequently absorbed, are beneficial to the rapidly growing embryonic calf by removing matter so precious as phosphate of lime? When a mans fingers have been amputated, imperfect nails have been known to appear on the stumps, and I could as soon believe that these vestiges of nails have been developed

rule, that a rudimentary part or organ is of greater size relatively to the adjoining parts in the embryo, than in the adult; so that the organ at this early age is less rudimentary, or even cannot be said to be in any degree rudimentary. Hence, also, a rudimentary organ in the adult, is often said to have retained its embryonic condition.

I have now given the leading facts with respect to rudimentary organs. In reflecting on them, every one must be struck with astonishment: for the same reasoning power which tells us plainly that most parts and organs are exquisitely adapted for certain purposes, tells us with equal plainness that these rudimentary or atrophied organs, are imperfect and useless. In works on natural history rudimentary organs are generally said to have been created "for the sake of symmetry," or in order "to complete the scheme of nature;" but this seems to me no explanation, merely a restatement of the fact. ↑ Would it be thought sufficient to say that because planets revolve in elliptic courses round the sun, satellites follow the same course round the planets, for the sake of symmetry, and to complete the scheme of nature? An eminent physiologist accounts for the presence of rudimentary organs, by supposing that they serve to excrete matter in excess, or injurious to the system; but can we suppose that the minute papilla, which often represents the pistil in male flowers, and which is formed merely of cellular tissue, can thus act? Can we suppose that the formation of rudimentary teeth which are subsequently absorbed, can be of any service to the rapidly growing embryonic calf by the excretion of precious phosphate of lime? When a man's fingers have been amputated, imperfect nails sometimes appear on the stumps: I could as soon believe that these vestiges of nails have appeared, not from unknown laws of growth, but