Origin of Species Variorum

Comparison with 1872

species constantly trying to increase in number, with natural selection always ready to adapt the slowly varying descendants of each to any unoccupied or ill-occupied place in nature, these facts cease to be strange, or .. might even have been anticipated.

We can understand how it is that such harmonious beauty generally prevails throughout nature. That there are exceptions according to our ideas of beauty, no one will doubt who will look at some of the venomous snakes, at some fish, and at certain hideous bats with a distorted resemblance to the human face. Sexual selection has given, generally to the males alone but sometimes to both sexes, the most brilliant and beautiful colours, as well as other ornaments, to our birds, butterflies, and a few other animals. It has rendered the voices of many male birds musical to their females, as well as to our ears. Flowers and fruit have been rendered conspicuous by gaudy colours in contrast with the green foliage, in order that the flowers might be easily seen, visited, and fertilised by insects, and the fruit have their seeds disseminated by birds. How it comes that certain colours, sounds, and forms should give pleasure to man and the lower animals,— that is, how the sense of beauty in its simplest form was first acquired,— we do not know any more than how certain odours and flavours were first rendered agreeable. ↑

As natural selection acts by competition, it adapts and improves the inhabitants of each country only in relation to their co-inhabitants; so that we need feel no surprise at the species of any one country, although on the ordinary view supposed to have been created and specially adapted for that country, being beaten and supplanted by the naturalised productions from another land. Nor ought we to marvel if all the contrivances in nature be not, as far as we can judge, absolutely perfect, as in the case even of the human eye; or if some of them be abhorrent to our ideas of fitness. We need not marvel at the sting of the bee, when used against an enemy, causing the bee's own death; at drones being produced in such great numbers for one single act, and being then slaughtered by their sterile sisters; at the astonishing waste of pollen by our fir-trees; at the instinctive hatred of the queen-bee for her own fertile daughters; at ichneumonidæ feeding within the living bodies of caterpillars; or at other such cases. The wonder indeed is, on the theory of natural selection, that more cases of the want of absolute perfection have not been detected.

The complex and little known laws governing the production of varieties are the same, as far as we can judge, with the laws which have governed the production of distinct species. In both cases physical conditions seem to have produced some direct and definite effect, but how much we cannot say. Thus when varieties enter any new station, they occasionally assume some of the characters proper to the species of that station. With both varieties and species, use and disuse seem to have produced a considerable effect; for it is impossible to resist this ..

As natural selection acts by competition, it adapts the inhabitants of each country only in relation to the degree of perfection of their associates; so that we need feel no surprise at the inhabitants of any one country, although on the ordinary view supposed to have been specially created and adapted for that country, being beaten and supplanted by the naturalised productions from another land. Nor ought we to marvel if all the contrivances in nature be not, as far as we can judge, absolutely perfect; and if some of them be abhorrent to our ideas of fitness. We need not marvel at the sting of the bee causing the bee's own death; at drones being produced in such vast numbers for one single act, and being then slaughtered by their sterile sisters; at the astonishing waste of pollen by our fir-trees; at the instinctive hatred of the queen bee for her own fertile daughters; at ichneumonidæ feeding within the live bodies of caterpillars; and at other such cases. The wonder indeed is, on the theory of natural selection, that more cases of the want of absolute perfection have not been observed.

The complex and little known laws governing variation are the same, as far as we can see, with the laws which have governed the production of so-called specific forms. In both cases physical conditions seem to have produced but little direct effect; yet when varieties enter any zone, they occasionally assume some of the characters of the species proper to that zone. In both varieties and species, use and disuse seem to have produced some effect; for it is difficult to resist this con-