Origin of Species Variorum

Comparison with 1859

the several species to their several places in the economy of nature, and likewise to fit the two sexes of the same species to each other, or to fit the males and females to different habits of life, or the males to struggle with other males for the possession of the females.

Finally, then, I conclude that the greater variability of specific characters, or those which distinguish species from species, than of generic characters, or those which the species possess in common;— that the frequent extreme variability of any part which is developed in a species in an extraordinary manner in comparison with the same part in its congeners; and the not great degree of variability in a part, however extraordinarily it may be developed, if it be common to a whole group of species;— that the great variability of secondary sexual characters, and the great amount of difference in these same characters between closely allied species;— that secondary sexual and ordinary specific differences are generally displayed in the same parts of the organisation,— are all principles closely connected together. All being mainly due to the species of the same group having descended from a common progenitor, from whom they have inherited much in common,— to parts which have recently and largely varied being more likely still to go on varying than parts which have long been inherited and have not varied,— to natural selection having more or less completely, according to the lapse of time, overmastered the tendency to reversion and to further variability,— to sexual selection being less rigid than ordinary selection,— and to variations in the same parts having been accumulated by natural and sexual selection, and thus adapted for secondary sexual, and for ordinary specific purposes.

Finally, then, I conclude that the greater variability of specific characters, or those which distinguish species from species, than of generic characters, or those which the species possess in common;— that the frequent extreme variability of any part which is developed in a species in an extraordinary manner in comparison with the same part in its congeners; and the slight degree of variability in a part, however extraordinarily it may be developed, if it be common to a whole group of species;— that the great variability of secondary sexual characters, and the great amount of difference in these same characters between closely allied species;— that secondary sexual and ordinary specific differences are generally displayed in the same parts of the organisation,— are all principles closely connected together. All being mainly due to the species of the same group having descended from a common progenitor, from whom they have inherited much in common,— to parts which have recently and largely varied being more likely still to go on varying than parts which have long been inherited and have not varied,— to natural selection having more or less completely, according to the lapse of time, overmastered the tendency to reversion and to further variability,— to sexual selection being less rigid than ordinary selection,— and to variations in the same parts having been accumulated by natural and sexual selection, and having been thus adapted for secondary sexual, and for ordinary specific purposes.