Origin of Species Variorum

a remarkable fact; more especially when we reflect how many species there are, which, though resembling each other most closely, are utterly sterile when intercrossed. Several considerations, however, render the fertility of domestic varieties less remarkable than at first appears. It can, in the first place, be clearly shown that mere external dissimilarity between two species does not determine their greater or lesser degree of sterility when crossed; and we may apply the same rule to domestic varieties. ↑ ↑ In the second place, some eminent naturalists believe that a long course of domestication tends to eliminate sterility in the successive generations of hybrids which were at first only slightly sterile; and if this be so, we surely ought not to expect to find sterility both appearing and disappearing under nearly the same conditions of life. ↑ Lastly, and this seems to me by far the most important consideration, new races of animals and plants are produced under domestication by man's methodical and unconscious power of selection, for his own use and pleasure: he neither wishes to select, nor could select, slight differences in the reproductive system, or other constitutional differences correlated with the reproductive system. ↑ He supplies his several varieties with the same food; treats them in nearly the same manner, and does not wish to alter their general habits of life. Nature acts uniformly and slowly during vast periods of time on the whole organisation, in any way which may be for each creature's own good; and thus she may, either directly, or more probably indirectly, through correlation, modify the reproductive system in the several descendants from any one species. Seeing this difference in the process of selection, as carried on by man and nature, we need not be surprised at some difference in the result.

I have as yet spoken as if the varieties of the same