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Comparison with 1866

breathing organ or lung. Other similar instances could be given.
Organs, however little developed, if of use, should not be called rudimentary: they may be called nascent, and may hereafter be developed by natural selection to any further extent. Rudimentary organs in the other hand, are essentially useless, as teeth which never cut through the gums. As they would be of even less use, when in a still less developed condition, they cannot have been formed through variation and natural selection, which latter acts solely by the preservation of useful modifications. They relate to a former state of things, and have been partially retained by the power of inheritance. It is difficult to know what organs are nascent; looking to the future, we cannot of course tell how any part will be developed, and whether it is now in a nascent condition; looking to the past, creatures with an organ in a nascent condition will generally have been supplanted .. by their successors with the same organ in a more perfect and developed condition, and consequently will not now exist. The wing of the penguin is of high service, acting as a fin; it may, therefore, represent the nascent state of the wing; .. not that I believe this to be the case; it is more probably a reduced organ, modified for a new function: the wing of the Apteryx, on the other hand, is quite useless, and is truly rudimentary. The simple filamentary limbs of the Lepidosiren apparently are in a nascent state; for, as Owen has remarked, they are the "beginnings of organs which attain full functional development in higher vertebrates." The mammary glands of the Ornithorhynchus may, probably, be considered, in comparison with those of the cow, as in a nascent condition. The ovigerous frena of certain cirripedes, which are only slightly developed and which have ceased to give attachment to the ova, are nascent branchiæ. Rudimentary organs in the individuals of the same species are very liable to vary in degree of development
breathing organ or lung. Other similar instances could be given.
Organs, however little developed, if of use, should not be called rudimentary; they cannot properly be said to be in an atrophied condition; they may be called nascent, and may hereafter be developed to any extent by natural selection. Rudimentary organs, on the other hand, are essentially useless, as teeth which never cut through the gums; in a still less developed condition, they would be of still less use. They cannot, therefore, under their present condition, have been formed by natural selection, which acts solely by the preservation of useful modifications; they have been retained, as we shall see, by inheritance, and relate to a former condition of their possessor. It is difficult to know what are nascent organs; looking to the future, we cannot of course tell how any part will be developed, and whether it is now nascent; looking to the past, creatures with an organ in a nascent condition will generally have been supplanted and exterminated by their successors with the organ in a more perfect and developed condition. The wing of the penguin is of high service, and acts as a fin; it may, therefore, represent the nascent state of the wings of birds; not that I believe this to be the case, it is more probably a reduced organ, modified for a new function: the wing of the Apteryx is useless, and is truly rudimentary. The mammary glands of the Ornithorhynchus may, perhaps, be considered, in comparison with the udder of a cow, as in a nascent state. The ovigerous frena of certain cirripedes, which are only slightly developed and which have ceased to give attachment to the ova, are nascent branchiæ. Rudimentary organs in the individuals of the same species are very liable to vary in degree of development