Origin of Species Variorum

Comparison with 1866

of the sea or of the land; and, consequently, the competition between fresh-water productions will have been less severe than elsewhere; new forms will have been more slowly formed, and old forms more slowly exterminated. And it is in fresh water that we find seven genera of Ganoid fishes, remnants of a once preponderant order: and in fresh water we find some of the most anomalous forms now known in the world, as the Ornithorhynchus and Lepidosiren, which, like fossils, connect to a certain extent orders now widely separated in the natural scale. These anomalous forms may .. be called living fossils; they have endured to the present day, from having inhabited a confined area, and thus having .. been exposed to less severe competition.

To sum up the circumstances favourable and unfavourable to natural selection, as far as the extreme intricacy of the subject permits. I conclude, looking to the future, that for terrestrial productions a large continental area, which will probably undergo many oscillations of level, and which consequently will exist for long periods in a broken condition, will be the most favourable for the production of many new forms of life, likely to endure long and to spread widely. For the area will first have existed as a continent, and the inhabitants, at this period numerous in individuals and kinds, will have been subjected to .. severe competition. When converted by subsidence into large separate islands, there will still exist many individuals of the same species on each island: intercrossing on the confines of the range of each species will thus be checked: after physical changes of any kind, immigration will be pre- vented, so that new places in the polity of each island will have to be filled up by modifications of the old inhabitants; and time will be allowed for the varieties in each to become well modified and perfected. When, by renewed elevation, the islands shall be re-converted into a continental area, there will again be severe competition: the most favoured or improved varieties will be enabled to spread: there will be much extinction of the less improved forms, and the relative proportional numbers of the various inhabitants of the renewed continent will again be changed; and again there will be a fair field for natural selection to improve still further the inhabitants, and thus produce new species.

That natural selection always acts with extreme slowness, I fully admit. Its action depends on there being places in the polity of nature, which can be better filled through some of the inhabitants of the country under- going modification of some kind. The existence of such places will often depend on physical changes, which are generally very slow, and on the immigration of better adapted forms having been checked. ↑ But the action of natural selection will probably still oftener depend on some of the inhabitants becoming slowly modified; the mutual relations of many of the other inhabitants being thus disturbed. Nothing can be effected, unless favourable variations occur, and variation itself is apparently always a very slow process. The process will often be greatly retarded by free intercrossing. Many will exclaim that these several causes are amply sufficient wholly to stop the action of natural selection. I do not believe so. On the other hand, I do believe that natural selection will always act very slowly, generally at only long intervals of time, and generally on only a very few of the inhabitants of the same region at the same time. I further believe, that this very slow,

of the sea or of the land; and, consequently, the competition between fresh-water productions will have been less severe than elsewhere; new forms will have been more slowly formed, and old forms more slowly exterminated. And it is in fresh water that we find seven genera of Ganoid fishes, remnants of a once preponderant order: and in fresh water we find some of the most anomalous forms now known in the world, as the Ornithorhynchus and Lepidosiren, which, like fossils, connect to a certain extent orders now widely separated in the natural scale. These anomalous forms may almost be called living fossils; they have endured to the present day, from having inhabited a confined area, and from having thus been exposed to less severe competition.

To sum up the circumstances favourable and unfavourable to natural selection, as far as the extreme intricacy of the subject permits. I conclude ... that for terrestrial productions a large continental area, which has undergone many oscillations of level, and which consequently has existed for long periods in a broken condition, has been the most favourable for the production of many new forms of life, fitted to endure long and to spread widely. For the area first existed as a continent, and the inhabitants, at this period numerous in individuals and kinds, will have been subjected to very severe competition. When converted by subsidence into large separate islands, there will still have existed many individuals of the same species on each island: intercrossing on the confines of the range of each species will thus have been checked: after physical changes of any kind, immigration will have been prevented, so that new places in the polity of each island will have had to be filled up by modifications of the old inhabitants; and time will have been allowed for the varieties in each to become well modified and perfected. When, by renewed elevation, the islands were reconverted into a continental area, there will again have been severe competition: the most favoured or improved varieties will have been enabled to spread: there will have been much extinction of the less improved forms, and the relative proportional numbers of the various inhabitants of the renewed continent will again have been changed; and again there will have been a fair field for natural selection to improve still further the inhabitants, and thus produce new species.

That natural selection will always act with extreme slowness I fully admit. Its action depends on there being places in the polity of nature, which can be better occupied by some of the inhabitants of the country undergoing modification of some kind. The existence of such places will often depend on physical changes, which are generally very slow, and on the immigration of better adapted forms having been checked. ↑ But the action of natural selection will probably still oftener depend on some few of the inhabitants becoming slowly modified; the mutual relations of many of the other inhabitants being thus disturbed. Nothing can be effected, unless favourable variations occur, and variation itself is apparently always a .. slow process. The process will often be greatly retarded by free intercrossing. Many will exclaim that these several causes are amply sufficient wholly to stop the action of natural selection. I do not believe so. On the other hand, I do believe that natural selection always acts very slowly, often only at long intervals of time, and generally on only a very few of the inhabitants of the same region at the same time. I further believe, that this slow,