Origin of Species Variorum

Comparison with 1869

relations of organic beings are more important; and as the number of species in any country goes on increasing, the organic conditions of life .. become more and more complex. Consequently there seems at first sight .. no limit to the amount of profitable diversification of structure, and therefore no limit to the number of species which might be produced. We do not know that even the most prolific area is fully stocked with specific forms: at the Cape of Good Hope and in Australia, which support such an astonishing number of species, many European plants have become naturalised. But geology shows us, that from an early part of the long tertiary period .. the number of species of shells, and that from the middle part of this same period the number of mammals, has not greatly or at all increased. What then checks an indefinite increase in the number of species? The amount of life (I do not mean the number of specific forms) supported on any area must have a limit, depending so largely as it does on physical conditions; therefore, if an area be inhabited by very many species, each or nearly each species will be represented by few individuals; and such species will be liable to extermination from accidental fluctuations in the nature of the seasons or in the number of their enemies. The process of extermination in these cases will be rapid, whereas the production of new species must always be sloW. Imagine the extreme case of as many species as individuals in England, and the first severe winter or very dry summer would exterminate thousands on thousands of species. Rare species, and each species will become rare if the number of species in any country becomes indefinitely increased, will, on the principle often explained, present within a given period few favourable variations; consequently, the process of giving birth to new specific forms will thus be retarded. When any species becomes very rare, close interbreeding will

relations of organic beings are more important; and as the number of species in any country goes on increasing, the organic conditions of life will become more and more complex. Consequently there seems at first sight to be no limit to the amount of profitable diversification of structure, and therefore no limit to the number of species which might be produced. We do not know that even the most prolific area is fully stocked with specific forms: at the Cape of Good Hope and in Australia, which support such an astonishing number of species, many European plants have become naturalised. But geology shows us, at least within the whole immense tertiary period, that the number of species of shells, and, probably, of mammals, has not greatly or at all increased. What then checks an indefinite increase in the number of species? The amount of life (I do not mean the number of specific forms) supported on any area must have a limit, depending so largely as it does on physical conditions: therefore, if an area be inhabited by very many species, each or nearly each species will be represented by few individuals; and such species will be liable to extermination from accidental fluctuations in the nature of the seasons or in the number of their enemies. The process of extermination in these cases will be rapid, whereas the production of new species will always be slow. Imagine the extreme case of as many species as individuals in England, and the first severe winter or very dry summer would exterminate thousands on thousands of species. Rare species, and each species will become rare if the number of species become in any country indefinitely increased, will, on the principle often explained, present within a given period few favourable variations; consequently, the process of giving birth to new specific forms will thus be retarded. When any species becomes very rare, close interbreeding will