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I do not 1859 1860 1861
it need not be 1866 1869 1872

the two little 1861 1866 1869 1872
little 1859 1860

8 blocks not present in 1859 1860 1861 1866 1869; present in 1872
There is another possible mode of transition, namely, through the acceleration or retardation of the period of reproduction. This has lately been insisted on by Prof. Cope and others in the United States. It is now known that some animals are capable of reproduction at a very early age, before they have acquired their perfect characters; and if this power became thoroughly well developed in a species, it seems probable that the adult stage of development would sooner or later be lost; and in this case, especially if the larva differed much from the mature form, the character of the species would be greatly changed and degraded. Again, not a few animals, after arriving at maturity, go on changing in character during nearly their whole lives. With mammals, for instance, the form of the skull is often much altered with age, of which Dr. Murie has given some striking instances with seals; every one knows how the horns of stags become more and more branched, and the plumes of some birds become more finely developed, as they grow older. Prof. Cope states that the teeth of certain lizards change much in shape with advancing years; with crustaceans not only many trivial, but some important parts assume a new character, as recorded by Fritz Müller, after maturity. In all such cases,— and many could be given,— if the age for reproduction were retarded, the character of the species, at least in its adult state, would be modified; nor is it improbable that the previous and earlier stages of development would in some cases be hurried through and finally lost. Whether species have often or ever been modified through this comparatively sudden mode of transition, I can form no opinion; but if this has occurred, it is probable that the differences between the young and the mature, and between the mature and the old, were primordially acquired by graduated steps.

←Subtitle not present 1859 1860 1861 Special Difficulties of the Theory of Natural Selection. 1872
it is 1859 1860 1861
for it is 1866 1872
for it is im- 1869

but, as Owen and others have 1859 1860 1861
As Owen has 1866 1869
OMIT 1872

intimate structure closely resembles that of common muscle; 1859 1860 1861
OMIT 1872
is much analogy between them and ordinary muscles, in their manner of action, in the influence on them of the nervous power and other stimulants such as strychnine, 1866
is much analogy between them and ordinary muscles, in their manner of action, in the influence on them of the nervous power and of stimulants such as strychnine, 1869

it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteucci asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. 1861
OMIT 1872
it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteuchi asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. 1859
it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteucei asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. 1860
some believe in their intimate structure. 1866 1869

2 blocks not present in 1859 1860 1861; present in 1866 1869 1872
We do not even in all cases know of what use these organs are; though in the Gymnotus and Torpedo they no doubt serve as powerful means of defence and perhaps for securing prey; yet in the Ray an analogous organ in the tail, even when greatly irritated, manifests, as lately observed by Matteucci, but little electricity; so little that it can hardly be of much use for these ends. Moreover, in the Ray, besides the organ just referred to, there is, as Dr. R. M'Donnell has shown, another organ near the head, not known to be electrical, but which apparently is the real homologue of the electric battery in the torpedo.

3 blocks not present in 1859 1860 1861 1866 1869; present in 1872
It is generally admitted that there exists between these organs and ordinary muscle a close analogy, in intimate structure, in the distribution of the nerves, and in the manner in which they are acted on by various reagents. It should, also, be especially observed that muscular contraction is accompanied by an electrical discharge; and, as Dr. Radcliffe insists, "in the electrical apparatus of the torpedo during rest, there would seem to be a charge in every respect like that which is met with in muscle and nerve during rest, and the discharge of the torpedo, instead of being peculiar, may be only another form of the discharge which attends upon the action of muscle and motor nerve." Beyond this we cannot at present go in the way of explanation; but as we know so little about the uses of these organs, and as we know nothing about the habits and structure of the progenitors of the existing electric fishes, it would be extremely bold to maintain that no serviceable transitions are possible by which these organs might have been gradually developed.

1 blocks not present in 1859 1860 1861 1872; present in 1866 1869
And lastly, as we know nothing about the lineal progenitors of any of these fishes, it must be admitted that we are too ignorant to be enabled to affirm that no transitions are possible, through which the electric organs might have been developed.

The electric organs 1859 1860 1861
These same organs at first appear to 1866 1869
These organs appear at first to 1872

which 1859 1860 1861
fish, of which 1866 1869 1872

of the other family; indeed, they graduate into each other. Therefore I do not
doubted
doubt
that the two little folds of skin, which originally served as ovigerous frena, but which, likewise, very slightly
aid in
aided
in the
the
act of respiration, have been gradually converted by natural selection into branchiæ, simply through an increase in their size and the obliteration of their adhesive glands. If all pedunculated cirripedes had become extinct, and they have
already
already
suffered far more extinction than have sessile cirripedes, who would ever have imagined that the branchiæ in this latter family had originally existed as organs for preventing the ova from being washed out of the sack?
Although we must be extremely cautious in concluding that any organ could not
possibly
possibly
have been produced by
successive,
successive
small, transitional
transitional
gradations,
yet
yet,
undoubtedly
undoubtedly,
serious
grave
cases of difficulty
occur.
occur,
some
some
of
of
which
which
will
will
be
be
discussed
discussed
in
in
my
my
future
future
work.
work.
One of the
most serious
gravest
is that of neuter insects, which are often
very
very
differently constructed from either the males or fertile females; but this case will be treated of in the next chapter. The electric organs of fishes offer another case of special difficulty; it is
possible
impossible
to conceive by what steps these wondrous organs have been
produced.
produced;
but, as Owen and others have
remarked,
remarked,
there
their
their
intimate structure closely resembles that of common muscle;
and
and
as
as
it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteucci asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible.
The electric organs offer another and
far
even
more serious difficulty; for they occur in
only
only
about a dozen
kinds
fishes,
of which several are widely remote in their affinities.
Generally
Generally
When
when
the same organ
is found
appears
in