of the other family; indeed, they graduate into each other. Therefore
I do not I do not 1859 1860 1861 |
it need not be 1866 1869 1872 |
doubt doubt 1859 1860 1861 | doubted 1866 1869 1872 |
that
the two little the two little 1861 1866 1869 1872 |
little 1859 1860 |
folds of skin, which originally served as ovigerous frena, but which, likewise, very slightly
aided aided 1859 1860 1861 1866 1872 | aid in 1869 |
the the 1859 1860 1861 1866 1869 | in the 1872 |
act of respiration, have been gradually converted by natural selection into branchiæ, simply through an increase in their size and the obliteration of their adhesive glands. If all pedunculated cirripedes had become extinct, and they have
already already 1859 1860 1861 1866 1869 | already 1872 |
suffered far more extinction than have sessile cirripedes, who would ever have imagined that the branchiæ in this latter family had originally existed as organs for preventing the ova from being washed out of the sack? ↑8 blocks not present in 1859 1860 1861 1866 1869; present in 1872 | There is another possible mode of transition, namely, through the acceleration or retardation of the period of reproduction.
This has lately been insisted on by Prof. Cope and others in the United States.
It is now known that some animals are capable of reproduction at a very early age, before they have acquired their perfect characters; and if this power became thoroughly well developed in a species, it seems probable that the adult stage of development would sooner or later be lost; and in this case, especially if the larva differed much from the mature form, the character of the species would be greatly changed and degraded.
Again, not a few animals, after arriving at maturity, go on changing in character during nearly their whole lives.
With mammals, for instance, the form of the skull is often much altered with age, of which Dr. Murie has given some striking instances with seals; every one knows how the horns of stags become more and more branched, and the plumes of some birds become more finely developed, as they grow older.
Prof. Cope states that the teeth of certain lizards change much in shape with advancing years; with crustaceans not only many trivial, but some important parts assume a new character, as recorded by Fritz Müller, after maturity.
In all such cases,— and many could be given,— if the age for reproduction were retarded, the character of the species, at least in its adult state, would be modified; nor is it improbable that the previous and earlier stages of development would in some cases be hurried through and finally lost.
Whether species have often or ever been modified through this comparatively sudden mode of transition, I can form no opinion; but if this has occurred, it is probable that the differences between the young and the mature, and between the mature and the old, were primordially acquired by graduated steps.
|
↑Subtitle not present 1859 1860 1861 |
Special Difficulties of
the
Theory
of
Natural
Selection. 1872 |
|
Although we must be extremely cautious in concluding that any organ could not
possibly possibly 1859 1860 1861 | possibly 1866 1869 1872 |
have been produced by
successive successive 1859 1860 1861 1866 1869 | successive, 1872 |
transitional transitional 1859 1860 1861 1866 1869 | small, transitional 1872 |
gradations,
yet, yet, 1859 1860 1861 | yet 1866 1869 1872 |
undoubtedly, undoubtedly, 1859 1860 1861 | undoubtedly 1866 1869 1872 |
grave grave 1859 1860 1861 | serious 1866 1869 1872 |
cases of difficulty
occur, occur, 1859 1860 1861 1866 1869 | occur. 1872 |
some some 1859 1860 1861 1866 1869 | some 1872 |
of of 1859 1860 1861 1866 1869 | of 1872 |
which which 1859 1860 1861 1866 1869 | which 1872 |
will will 1859 1860 1861 1866 1869 | will 1872 |
be be 1859 1860 1861 1866 1869 | be 1872 |
discussed discussed 1859 1860 1861 1866 1869 | discussed 1872 |
in in 1859 1860 1861 1866 1869 | in 1872 |
my my 1859 1860 1861 1866 1869 | my 1872 |
future future 1859 1860 1861 1866 1869 | future 1872 |
work. work. 1859 1860 1861 1866 1869 | work. 1872 |
|
|
One of the
gravest gravest 1859 1860 1861 | most serious 1866 1869 1872 |
is that of neuter insects, which are often
very very 1859 1860 1861 | very 1866 1869 1872 |
differently constructed from either the males or fertile females; but this case will be treated of in the next chapter. The electric organs of fishes offer another case of special difficulty;
it is it is 1859 1860 1861 |
for it is 1866 1872 |
for it is im- 1869 |
impossible impossible 1859 1860 1861 1866 1872 | possible 1869 |
to conceive by what steps these wondrous organs have been
produced; produced; 1859 1860 1861 | produced. 1866 1869 1872 |
but, as Owen and others have but, as Owen and others have 1859 1860 1861 |
As Owen has 1866 1869 |
OMIT 1872 |
remarked, remarked, 1859 1860 1861 1866 1869 | remarked, 1872 |
their their 1859 1860 1861 | there 1866 1869 | their 1872 |
intimate structure closely resembles that of common muscle; intimate structure closely resembles that of common muscle; 1859 1860 1861 |
OMIT 1872 |
is much analogy between them and ordinary muscles, in their manner of action, in the influence on them of the nervous power and other stimulants such as strychnine, 1866 |
is much analogy between them and ordinary muscles, in their manner of action, in the influence on them of the nervous power and of stimulants such as strychnine, 1869 |
and and 1859 1860 1861 1866 1869 | and 1872 |
as as 1859 1860 1861 1866 1869 | as 1872 |
it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteucci asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteucci asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. 1861 |
OMIT 1872 |
it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteuchi asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. 1859 |
it has lately been shown that Rays have an organ closely analogous to the electric apparatus, and yet do not, as Matteucei asserts, discharge any electricity, we must own that we are far too ignorant to argue that no transition of any kind is possible. 1860 |
some believe in their intimate structure. 1866 1869 |
↑2 blocks not present in 1859 1860 1861; present in 1866 1869 1872 | We
do not even in all cases
know of what use these organs are; though in
the Gymnotus and Torpedo they no doubt serve as powerful means of defence
and perhaps for securing prey; yet in the Ray
an
analogous organ in the tail,
even when
greatly irritated,
manifests, as lately observed by Matteucci, but little electricity;
so little
that it can hardly be of much
use for these ends.
Moreover, in the Ray, besides the organ just referred to, there is, as Dr. R. M'Donnell has shown, another organ near the head, not known to be electrical, but which apparently is
the real homologue of the electric battery in the torpedo.
|
↑3 blocks not present in 1859 1860 1861 1866 1869; present in 1872 | It is generally admitted that there exists between these organs and ordinary muscle a close analogy, in intimate structure, in the distribution of the nerves, and in the manner in which they are acted on by various reagents.
It should, also, be especially observed that muscular contraction is accompanied by an electrical discharge; and, as Dr. Radcliffe insists, "in the electrical apparatus of the torpedo during rest, there would seem to be a charge in every respect like that which is met with in muscle and nerve during rest, and the discharge of the torpedo, instead of being peculiar, may be only another form of the discharge which attends upon the action of muscle and motor nerve."
Beyond this we cannot at present go in the way of explanation; but as we know so little about the uses of these organs, and as we know nothing about the habits and structure of the progenitors of the existing electric fishes, it would be extremely bold to maintain that no serviceable transitions are possible by which these organs might have been gradually developed.
|
↑1 blocks not present in 1859 1860 1861 1872; present in 1866 1869 | And
lastly,
as we know nothing about the lineal progenitors of any of
these fishes, it must be admitted that we are too ignorant to be enabled to affirm that no transitions are possible, through which the electric organs might have been developed.
|
|
|
The electric organs The electric organs 1859 1860 1861 |
These same organs at first appear to 1866 1869 |
These organs appear at first to 1872 |
offer another and
even even 1859 1860 1861 | far 1866 1869 1872 |
more serious difficulty; for they occur in
only only 1859 1860 1861 | only 1866 1869 1872 |
about a dozen
fishes, fishes, 1859 1860 1861 | kinds 1866 1869 1872 |
of
which which 1859 1860 1861 |
fish, of which 1866 1869 1872 |
several are widely remote in their affinities.
Generally Generally 1859 1860 1861 1866 1869 | Generally 1872 |
when when 1859 1860 1861 1866 1869 | When 1872 |
the same organ
appears appears 1859 1860 1861 | is found 1866 1869 1872 |
in
|