Origin of Species Variorum

are so poorly developed, that no record of several successive and peculiar marine faunas will probably be preserved to a distant age. A little reflection will explain why along the rising coast of the western side of South America, no extensive formations with recent or tertiary remains can anywhere be found, though the supply of sediment must for ages have been great, from the enormous degradation of the coast-rocks and from muddy streams entering the sea. The explanation, no doubt, is, that the littoral and sub-littoral deposits are continually worn away, as soon as they are brought up by the slow and gradual rising of the land within the grinding action of the coast-waves.

We may, I think, safely conclude that sediment must be accumulated in extremely thick, solid, or extensive masses, in order to withstand the incessant action of the waves, when first upraised and during subsequent oscillations of level. Such thick and extensive accumulations of sediment may be formed in two ways; either in profound depths of the sea, in which case, judging from the researches of E. Forbes , we may conclude that the bottom will be inhabited by few animals, but it will not be, as we at last know from the telegraphic soundings, barren of life; consequently the mass when up-raised will give a most imperfect record of the forms of life which existed during the period of deposition. Or, sediment may be accumulated to any thickness and extent over a shallow bottom, if it continue slowly to subside. In this latter case, as long as the rate of subsidence and supply of sediment nearly balance each other, the sea will remain shallow and favourable for life, and thus a rich fossiliferous formation thick enough, when upraised, to resist any amount of degradation, may be formed.

I am convinced that nearly all our ancient formations, which