Origin of Species Variorum

Comparison with 1866

ranked by all naturalists as closely allied to, but specifically distinct from, their northern representatives.

Now let us see what light can be thrown on the foregoing facts, on the belief, supported as it is by a large body of geological evidence, that the whole world, or a large part of it, was simultaneously colder during the Glacial period than at present. The Glacial period, as measured by years, must have been very long; and when we remember over what vast spaces some naturalised plants and animals have spread within a few centuries, this period will have been ample for any amount of migration. As the cold came slowly on, the tropical plants and animals will have retreated from both sides towards the equator, followed in the rear by the temperate productions, and these by the arctic; but with the latter we are not now concerned. The whole problem of what will have occurred is excessively complex. The probable existence before the Glacial period of a pleistocene equatorial flora and fauna, fitted for a hotter climate than any now existing, must not be over-looked. This old equatorial flora will have been almost wholly destroyed, and the two pleistocene sub-tropical floras, commingled and reduced in number, will then have formed the equatorial flora. There will also probably have been during the Glacial period great changes in the precise nature of the climate, in the degree of humidity, &c.; and various animals and plants will have migrated in different proportions and at different rates. So that altogether during the Glacial period the inhabitants of the tropics must have been greatly disturbed in all their relations of life. ↑ Hence they will have suffered much extinction; how much no one can say; perhaps formerly the tropics supported as many species as we now see crowded together at the ... Cape of Good Hope, and in parts of temperate Australia. As we know that many tropical plants and animals can withstand a considerable amount of cold, many might have escaped extermination during a moderate fall of temperature, more especially by escaping into the warmest spots. Nor must it be overlooked that, as the cold will have come on very slowly, it is almost certain that many of the inhabitants of the tropics will have become in some degree acclimatised; in the same manner as the same species of plant when living on lowlands and highlands certainly transmit to their seedlings different constitutional powers of resisting cold. Nevertheless, it cannot be denied that all tropical productions will have greatly suffered, and the chief difficulty is to understand how they can have escaped entire annihilation. ↑ On the other hand, the temperate productions, after migrating nearer to the equator, though they will have been placed under somewhat new conditions, will have suffered less. And it is certain that many temperate plants, if protected from

ranked by some naturalists as specifically distinct, by others as varieties; but some are certainly identical, and many, though closely related to northern forms, must be ranked as distinct species.

Now let us see what light can be thrown on the foregoing facts, on the belief, supported as it is by a large body of geological evidence, that the whole world, or a large part of it, was during the Glacial period simultaneously much colder than at present. The Glacial period, as measured by years, must have been very long; and when we remember over what vast spaces some naturalised plants and animals have spread within a few centuries, this period will have been ample for any amount of migration. As the cold came slowly on, all the tropical plants and other productions will have retreated from both sides towards the equator, followed in the rear by the temperate productions, and these by the arctic; but with the latter we are not now concerned. ↑ ↑ The tropical plants probably suffered much extinction; how much no one can say; perhaps formerly the tropics supported as many species as we see at the present day crowded together at the Cape of Good Hope, and in parts of temperate Australia. As we know that many tropical plants and animals can withstand a considerable amount of cold, many might have escaped extermination during a moderate fall of temperature, more especially by escaping into the lowest, most protected, and warmest districts. ↑ But the great fact to bear in mind is, that all tropical productions will have suffered to a certain extent. On the other hand, the temperate productions, after migrating nearer to the equator, though they will have been placed under somewhat new conditions, will have suffered less. And it is certain that many temperate plants, if protected from