Origin of Species Variorum

Comparison with 1869

The different orders of insects inhabiting Madeira .. present nearly similar cases.

Oceanic islands are sometimes deficient in animals of certain whole classes, and their places are .. occupied by other classes: thus in the Galapagos Islands reptiles, and in New Zealand gigantic wingless birds, take, or recently took, the place of mammals. Although New Zealand is here spoken of as an oceanic island, it is in some degree doubtful whether it should be so ranked; it is of large size, and is not separated from Australia by a profoundly deep sea: from its geological character and the direction of its mountain-ranges, the Rev. W. B. Clarke has lately maintained that this island, as well as New Caledonia, should be considered as appurtenances of Australia. Turning to plants, Dr. Hooker has shown that in the Galapagos Islands the proportional numbers of the different orders are very different from what they are elsewhere. All such differences in number, and the absence of certain whole groups of animals and plants on islands, are generally accounted for by supposed differences in their physical conditions; ... but this explanation is not a little doubtful. Facility of immigration, seems to have been fully as important as the nature of the conditions.

Many remarkable little facts could be given with respect to the inhabitants of oceanic islands. For instance, in certain islands not tenanted by a single mammal, some of the endemic plants have beautifully hooked seeds; and few relations are more manifest than that hooked seeds are adapted for transportal in the wool or fur of quadrupeds. But a hooked seed might be carried to an island by .. other means; and the plant then becoming .. modified ... would form an endemic species, which might still retain the hooks, which would not form a more useless appendage than the shrivelled wings under the soldered elytra of many insular beetles. Again, islands often possess trees or bushes belonging to orders .. elsewhere including only herbaceous species; now trees, as Alph. de Candolle has shown, generally have, what- ever the cause may be, confined ranges. Hence trees would he little likely to reach distant oceanic islands; and an herbaceous plant, which had no chance of successfully

The different orders of insects in Madeira apparently present analogous facts.

Oceanic islands are sometimes deficient in certain classes, and their places are apparently occupied by the other inhabitants; in the Galapagos Islands reptiles, and in New Zealand gigantic wingless birds, take the place of mammals. ↑ In the plants of the Galapagos Islands, Dr. Hooker has shown that the proportional numbers of the different orders are very different from what they are elsewhere. Such cases are generally accounted for by the physical conditions of the islands; but this explanation seems to me not a little doubtful. Facility of immigration seems to have been at least as important as the nature of the conditions.

Many remarkable little facts could be given with respect to the inhabitants of remote islands. For instance, in certain islands not tenanted by mammals, some of the endemic plants have beautifully hooked seeds; yet few relations are more striking than the adaptation of hooked seeds for transportal by the wool and fur of quadrupeds. This case presents no difficulty on my view, for a hooked seed might be transported to an island by some other means; and the plant then becoming slightly modified, but still retaining its hooked seeds, would form an endemic species, having as useless an appendage as any rudimentary organ,— for instance, as the shrivelled wings under the soldered elytra of many insular beetles. Again, islands often possess trees or bushes belonging to orders which elsewhere include only herbaceous species; now trees, as Alph. de Candolle has shown, generally have, whatever the cause may be, confined ranges. Hence trees would be little likely to reach distant oceanic islands; and an herbaceous plant, though it would have no chance of suc- cessfully