Origin of Species Variorum

Comparison with 1869

inhabitants of each area are related to the inhabitants of the nearest source whence immigrants might have been derived. We see this in the striking relation of nearly all the plants and animals of the Galapagos archipelago, of Juan Fernandez, and of the other American islands, ... to the plants and animals of the neighbouring American mainland; and of those of the Cape de Verde archipelago and of the other African islands to the African mainland. It must be admitted that these facts receive no explanation on the theory of creation.

The fact, as we have seen, that all past and present organic beings can be arranged within a few great classes, in groups subordinate to groups, and with the extinct groups often falling in between the recent groups, is intelligible on the theory of natural selection with its contingencies of extinction and divergence of character. On these same principles we see how it is, that the mutual affinities of the forms within each class are so complex and circuitous. We see why certain characters are far more serviceable than others for classification;— why adaptive characters, though of paramount importance to the beings, are of hardly any importance in classification; why characters derived from rudimentary parts, though of no service to the beings, are often of high classificatory value; and why embryological characters are often the most valuable of all. The real affinities of all organic beings, in contradistinction to their adaptive resemblances, are due to inheritance or community of descent. The Natural System is a genealogical arrangement, with the acquired grades of dif- ference, marked by the terms, varieties, species, genera, families, &c.; and we have to discover the lines of descent by the most permanent characters whatever they may be and of however slight vital importance. .. ..

The similar framework of bones in the hand of a man, wing of a bat, fin of the porpoise, and leg

inhabitants of each area are related to the inhabitants of the nearest source whence immigrants might have been derived. We see this in nearly all the plants and animals of the Galapagos archipelago, of Juan Fernandez, and of the other American islands being related in the most striking manner to the plants and animals of the neighbouring American mainland; and those of the Cape de Verde archipelago and other African islands to the African mainland. It must be admitted that these facts receive no explanation on the theory of creation.

The fact, as we have seen, that all past and present organic beings constitute one grand natural system, with group subordinate to group, and with extinct groups often falling in between recent groups, is intelligible on the theory of natural selection with its contingencies of extinction and divergence of character. On these same principles we see how it is, that the mutual affinities of the species and genera within each class are so complex and circuitous. We see why certain characters are far more serviceable than others for classification;— why adaptive characters, though of paramount importance to the being, are of hardly any importance in classification; why characters derived from rudimentary parts, though of no service to the being, are often of high classificatory value; and why embryological characters are the most valuable of all. The real affinities of all organic beings are due to inheritance or community of descent. The natural system is a genealogical arrangement, in which we have to discover the lines of descent by the most permanent characters, however slight their vital importance may be.

The framework of bones being similar in the hand of a man, wing of a bat, fin of the porpoise, and leg