Origin of Species Variorum

Comparison with 1869

districts will, also, be prevented. Moritz Wagner has lately published an interesting essay on this subject, and has shown that the service rendered by isolation in preventing crosses between newly formed varieties is probably greater even than I have supposed. But from reasons already assigned I can by no means agree with this naturalist, that migration and isolation are necessary for the formation of new species. The importance of isolation is likewise great in preventing, after any physical change in the conditions, such as of climate, elevation of the land, &c., the immigration of better adapted organisms; and thus new places in the natural economy of the district are left open for the old inhabitants to struggle for and become adapted to. ↑ Lastly, isolation ... will give time for a new variety to be slowly improved; and this may sometimes be of importance in the production of new species. If, however, an isolated area be very small, either from being surrounded by barriers, or from having very peculiar physical conditions, the total number of the inhabitants will be small; and this will retard the production of new species through natural selection, by decreasing the chances of the appearance of favourable individual differences.

The mere lapse of time by itself does nothing either for or against natural selection. I state this because it has been erroneously asserted that the element of time has been assumed by me to play an all-important part in modifying species, as if all ... were necessarily under- going change through the action of some innate law. Lapse of time is only so far .. important, and its importance in this respect is great, that it gives a better chance of beneficial variations arising, being selected, increased, and fixed, in relation to the slowly changing organic and inorganic conditions of life. It likewise favours the definite action of the conditions of life.

If we turn to nature to test the truth of these remarks, and look at any small isolated area, such as an oceanic island, although the .. number of the different species inhabiting it is small, as we shall see

districts, will be prevented. ↑ But isolation probably acts more efficiently in checking the immigration of better adapted organisms, after any physical change, such as of climate or elevation of the land, &c.; and thus new places in the natural economy of the country are left open for the old inhabitants to struggle for and become adapted to, through modifications in their structure and constitution. Lastly, isolation, by checking immigration and consequently competition, will give time for any new variety to be slowly improved; and this may sometimes be of importance in the production of new species. If, however, an isolated area be very small, either from being surrounded by barriers, or from having very peculiar physical conditions, the total number of the individuals supported on it will necessarily be very small; and fewness of individuals will greatly retard the production of new species through natural selection, by decreasing the chance of the appearance of favourable variations.

The mere lapse of time by itself does nothing either for or against natural selection. I state this because it has been erroneously asserted that the element of time is assumed by me to play an all-important part in natural selection, as if all species were necessarily undergoing slow modification from some innate law. Lapse of time is only so far highly important, as it gives a better chance of beneficial variations arising, being selected, accumulated, and fixed, in relation to the slowly changing organic and inorganic conditions of life. It likewise favours the direct action of new or changed physical conditions of life.

If we turn to nature to test the truth of these remarks, and look at any small isolated area, such as an oceanic island, although the total number of the species inhabiting it will be found to be small, as we shall see